Metaphalangium cirtanum, copyright Luigi Barraco.

Belongs within: Phalangiidae.

Metaphalangium is a genus of harvestmen with a distribution centred in the Mediterranean region. Members of this genus have a sandy yellow dorsum with a darker saddle and a pale median stripe, and males have a broad and deep subapical spoon on the penis (Snegovaya & Staręga 2009).

Characters (from Staręga 1984): Body coloration with saddle in middle part of abdomen angularly enlarged (sometimes only this portion of the saddle visible), and a clear white longitudinal band from eye-mound to anus. Cephalothorax denticulate anterior to eye-mound, tergites with transverse rows of larger, sharper denticles. Eye-mound normal, far from anterior edge of carapace (up to twice its length), with long denticles on the eye-rings. Supracheliceral lamella with distinct granules or denticles, sometimes forming a transverse series. Chelicerae of male usually more developed than those of female and with several denticles or granules. Pedipalps short and strong, femur usually with strong denticles, patella without apophysis, tarsus in male with ventral strip of sensory granules. Legs long and strong, at least the femora with longitudinal rows of strong denticles. Leg I (femur to tibia) greatly thickened in male. Penis with characteristic subapical “spoon”, sometimes occupying almost half length of shaft. Truncus expanded basally, sometimes in form of pedestal. Glans relatively large, triangular in profile, wedge-shaped in cross-section. Stylus relatively long.

<==Metaphalangium Roewer 1911K14 (see below for synonymy)
    |--M. cirtanum (Koch 1839)K14 (see below for synonymy)
    |--M. abruptum (Roewer 1911)SS09 (see below for synonymy)
    |--M. albiunilineatum (Lucas 1846)SS09 (see below for synonymy)
    |--M. bispinifrons (Roewer 1911) (see below for synonymy)S84
    |--M. corsicum (Roewer 1956)SS09
    |--M. lusitanicum (Roewer 1956)SS09
    |--M. monticola (Mkheidze 1952) [=Paropilio monticola]KM20
    |--M. orientale Staręga 1973S84
    |--M. punctatum (Roewer 1956) [=Paropilio punctatus]KM20
    |--M. spinipes (Roewer 1956) [=Paropilio spinipes]KM20
    |--M. strandi (Nosek 1905)K14 [=Phalangium strandiR23, Paropilio strandiR23]
    |--M. sudanum Roewer 1961S84
    `--M. tuberculatum (Lucas 1846)SS09 (see below for synonymy)

Metaphalangium Roewer 1911K14 [incl. Apozacheus Šilhavý 1965C92, Metegaenus Šilhavý 1965C92, Odontosoma Šilhavý 1946 non Townsend 1916C92, Parodontosoma Šilhavý 1965C92, Paropilio Roewer 1911C92]

Metaphalangium abruptum (Roewer 1911)SS09 [=Phalangium abruptumR23, Paropilio abruptusR23; incl. Pa. abstrusus Roewer 1911 non Ph. abstrusum Koch 1882R23]

Metaphalangium albiunilineatum (Lucas 1846)SS09 [=Phalangium albounilineatumS84; incl. Lacinius denticulatus Roewer 1923C92, *Parodontosoma denticulatumC92]

Metaphalangium bispinifrons (Roewer 1911) [=Zacheus bispinifrons; incl. Z. maroccanus Roewer 1915, Metaphalangium maroccanum Roewer 1923 non Zacheus maroccanus Roewer 1915]S84

Metaphalangium cirtanum (Koch 1839)K14 [=Opilio cirtanusD04, Phalangium cirtanumS84; incl. Ph. abstrusum Koch 1882C92, Metaphalangium abstrusumPWF22, *Paropilio abstrususC92, Phalangium africanum Lucas 1847D04, Metadasylobus albanicus Schenkel 1947D04, Ph. annulipes Lucas 1847D04, Lacinius armatus Roewer 1923D04, Ph. barbarum Lucas 1847D04, *Odontosoma centetes Šilhavý 1946C92, Ph. clavipus Roewer 1911PWF22, Metaphalangium propinquum denticulatum Hadži 1973D04, Ph. echinatum Lucas 1847D04, Acantholophus echinatus nec Boisduval 1835 nec Guérin-Méneville 1838 (both not preoc. as in homonymous gen.)D04, Lacinius echinatusS84, Phalangium infuscatum Lucas 1847D04, Dasylobus infuscatusR23, Eudasylobus infuscatusD04, Phalangium instabile Lucas 1847D04, Opilio luridus Koch 1848D04, Zacheus moesiacus Roewer 1923D04, *Apozacheus moesiacusC92, Metaphalangium propinquum montenegrinum Hadži 1973D04, Phalangium numidicum Lucas 1847D04, Ph. propinquum Lucas 1846SS09, *Metaphalangium propinquumC92, M. propinquum romanum Roewer 1923D04, Phalangium semiechinatum Simon 1885D04, Paropilio semiechinatusS84]

Metaphalangium tuberculatum (Lucas 1846)SS09 [=Phalangium tuberculatumD04, Egaenus tuberculatusS84, Egoenus (l. c.) tuberculatusS84, Lacinius tuberculatusD04, *Metegaenus tuberculatusC92, Oligolophus tuberculatusD04; incl. Dacnopilio carthaginiensis Roewer 1923D04, Dac. carthaginensis (l. c.)S84, Egaenus kabylicus Roewer 1956D04, Dasylobus serrifer Simon 1878D04, Eudasylobus serriferS84]

*Type species of generic name indicated


[C92] Crawford, R. L. 1992. Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History 8: 1–60.

[D04] Delfosse, E. 2004. Catalogue préliminaire des Opilions de France métropolitaine (Arachnida Opiliones). Bulletin de Phyllie 20: 34–58.

[K14] Kurt, K. 2014. Updated checklist of harvestmen (Arachnida: Opiliones) in Turkey. Arch. Biol. Sci. 66 (4): 1617–1631.

[KM20] Kury, A. B., A. C. Mendes, L. Cardoso, M. S. Kury & A. A. Granado. 2020. WCO-Lite: Online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0—Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018. Published by the author: Rio de Janeiro.

[PWF22] Prieto, C. E., H. Wijnhoven & J. B. Febrer. 2022. New harvestmen (Arachnida: Opiliones) from Menorca Island. Revista Ibérica de Aracnología 40: 145–157.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

[SS09] Snegovaya, N. Y., & W. Staręga. 2009. Taurolaena, a new genus of Phalangiidae (Opiliones). Revista Ibérica de Aracnología 17: 37–44.

[S84] Staręga, W. 1984. Revision der Phalangiidae (Opiliones), III. Die afrikanischen Gattungen der Phalangiinae, nebst Katalog aller afrikanischen Arten der Familie. Annales Zoologici 38 (1): 1–79.

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