Phalacrotophora epeirae, copyright John R. Maxwell.

Belongs within: Phoridae.
Contains: Melaloncha, Megaselia.

The Metopininae are a diverse group of scuttle flies that generally lack strong isolated bristles on the tibiae. Members of the genus Beckerina have two or four strongly divergent supra-antennal bristles that are usually more or less reclinate and porrect; in other metopinines, these bristles (if present) are not strongly divergent and usually proclinate (Petersen 1987).

Characters (from Petersen 1987): Frons sometimes bare along margins or on anteroventral half; supra-antennal bristles variable. Tibiae usually without bristles (exceptionally with a single anterodorsal bristle very near base on mid and hind tibiae) or with rows of strong setae. Wing generally fully developed, but sometimes reduced, rudimentary, or absent in female of some species.

| i. s.: Melittophora saltiB04
| MelalonchaB04
| Sinophoritis Hong 2002H02
| `--*S. bikerkosa Hong 2002H02
| Latiarista Hong 2002H02
| `--*L. intermixta Hong 2002H02
|--Beckerina [Beckerinini]P87
| |--B. alienaP87
| `--B. umbrimargoP87
| |--MegaseliaB10
| `--GymnophoraB10
| |--G. arcuataB10
| |--G. luteiventrisP87
| `--G. spiracularisB10
|--Ecitomyia wheeleriB10, P87
|--Rhabdomyia setosaB10
|--Lecanocerus compressicepsP87, B10
|--*Syneura cocciphilaB10, P87
|--Woodiphora magnipalpisP87
|--Cremersia spinicaudaP87, B10
|--Zyziphora hirtifronsP87
|--Myrmosicarius texanusP87, B10
|--Auxanommatidia californicaP87
|--Physoptera vesiculataP87, B10
|--Trophithauma rostratumP87, B10
|--Cataclinusa pachycondylaeP87, B10
|--Stenophorina petiolataP87
|--Chonocephalus americanusP87
|--Xanionotum mexicanumP87
|--Commoptera solenopsidisP87, B10
| |--A. brasiliensisP87
| `--A. melanderiP87
| |--D. appretiataB10
| `--D. disparifronsP87
| |--P. cataP87
| `--P. floricolaP87
| |--P. formicarumA71
| `--P. onyxP87
| |--T. furcatus [=Bactropalpus furcatus]B10
| `--T. pygmaeusB10
|--Metopina [incl. Typhlophorina]B10
| |--M. goeleti Grimaldi 1989RJ93
| |--M. pachycondylaeA71
| `--M. subarcuataB10
| |--P. borinquenensisB10
| |--P. imbecillaB10
| `--P. occidentalisP87
| |--A. asymmetricusB10
| |--A. (Mesophora) borealisP87
| `--A. wheeleriB10
| |--R. conicaP87
| |--R. maculineuraB10
| `--R. trivittataB10
| |--N. arnaudiP87
| |--N. cognataB10
| |--N. curvinervisB10
| `--N. mexicanaeB10
|--P. berolinensisB04
|--P. beukiB04
|--P. epeiraeB04
|--P. fasciataB04
|--P. halictorumB04
|--P. (Omapanta) longifronsP87
|--P. quadripunctataB04
`--P. rufiventrisB04

*Type species of generic name indicated


[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B04] Brown, B. V. 2004. Revision of the subgenus Udamochiras of Melaloncha bee-killing flies (Diptera: Phoridae: Metopininae). Zoological Journal of the Linnean Society 140: 1–42.

[B10] Brown, B. V. 2010. Phoridae (hump-backed flies, scuttle flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 725–761. NRC Research Press: Ottawa.

[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.

[P87] Peterson, B. V. 1987. Phoridae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 689–712. Research Branch, Agriculture Canada.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

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