Mitobatinae

Neoancistrotus gracilis, copyright Osvaldo Villarreal Manzanilla.

Belongs within: Gonyleptidae.

The Mitobatinae are a group of gonyleptid harvestmen with a usually elongate, unarmed femur IV in the male (Bragagnolo & Pinto-da-Rocha 2012).

Characters (from Bragagnolo & Pinto-da-Rocha 2012): Anterior margin of prosoma with frontal hump present and unarmed. Armature of ocularium usually with two spiniform apophyses. Armature of scutal areas variable, area III usually armed with pair of spiniform apophyses. Usually four areas on dorsal scutum; sometimes areas III and IV fused. Posterior margin may be armed with pair of tubercles. Free tergites unarmed or with median tubercles. Chelicera of similar size in both sexes. Femur of pedipalps only with short and thick ventral apical setae, patella and tibia unarmed. Coxa IV dorsally visible in entire extension in both sexes, with dorsoapical apophysis that can be robust and bifid in males (sexual dimorphism) or extremely reduced. Male femur IV straight and usually much elongated, usually more than four times scutal size, rarely about three times, and usually unarmed. Penis with ventral plate well defined, usually pyriform (sometimes rectangular), with setae on distal margin. Ventral process present; dorsal process absent. Stylus elongate, usually straight.

<==Mitobatinae
    |--Metamitobates Roewer 1913 (see below for synonymy)K03
    |    `--M. squalidus (Perty 1833)MKM22 (see below for synonymy)
    |--Despirus Roewer 1929 [incl. Ibarroides Mello-Leitão 1932, Segadasius Mello-Leitão 1949]K03
    |    `--*D. parvulus (Roewer 1927) (see below for synonymy)K03
    |--Dyscocyrtoides Mello-Leitão 1923KA-Z11 [=DiscocyrtoidesK03; incl. Mitobatoides Mello-Leitão 1927K03]
    |    `--D. nigricans (Mello-Leitão 1922) (see below for synonymy)K03
    |--Mitobatula Roewer 1931K03
    |    `--*M. castanea Roewer 1931K03
    |--Encheiridium Kury 2003K03
    |    |--*E. ruschii (Mello-Leitão 1942) [=Despirus ruschii, Discocyrtoides ruschii]K03
    |    `--E. montanum (Mello-Leitão 1941) [=Despirus montanus, Discocyrtoides montanus]K03
    |--Ischnotherus Kury 1991K03
    |    |--*I. tenebrosus Kury 1991K03
    |    `--I. pardus (Kury 1989) [=Discocyrtoides pardus]K03
    |--Ruschia Mello-Leitão 1940 non Schwantes 1926 (ICBN)K03
    |    |--*R. vellutina Mello-Leitão 1940 [=Discocyrtoides vellutinus]K03
    |    `--R. maculata (Soares 1974) [=Discocyrtoides maculatus]K03
    |--Mitobates Sundevall 1833K03 (see below for synonymy)
    |    |--*M. triangulus Sundevall 1833K03 (see below for synonymy)
    |    |--M. elegantulus (Mello-Leitão 1940) [=Neoancistrotus elegantulus, Ancistrotellus elegantulus]K03
    |    |--M. inermis Kury 1989K03
    |    `--M. pulcher (Mello-Leitão 1932) [=Despiroides pulcher, Discocyrtoides pulcher]K03
    |--Longiperna Roewer 1929K03
    |    |--*L. concolor (Mello-Leitão 1923) [=Discocyrtoides concolor]K03
    |    |--L. areolata (Soares 1944) [=Discocyrtoides areolatus]K03
    |    |--L. cancellata (Roewer 1913)DG09 [=Neomitobates cancellatusK03, Metamitobates cancellatusK03]
    |    |--L. coxalis (Roewer 1943) [=Mitobatula coxalis]K03
    |    |--L. curitibana Kury 2003 [=L. cancellata Roewer 1943 non Neomitobates cancellatus Roewer 1913]K03
    |    |--L. heliaca (Soares 1942) [=Despirus heliacus, Discocyrtoides heliacus]K03
    |    |--L. insperata (Soares & Soares 1947) [=Ancistrotellus insperatus]K03
    |    |--L. paranensis (Soares & Soares 1947) [=Discocyrtoides paranensis]K03
    |    `--L. zonata (Mello-Leitão 1935) [=Despirus zonatus, Discocyrtoides zonatus]K03
    |--Promitobates Roewer 1913K03 (see below for synonymy)
    |    |--*P. hexacanthus (Koch 1839)K03, DG09 [=Ancistrotus hexacanthusK03, Goniosoma hexacanthumK03]
    |    |--P. bellus (Soares 1945) [=Ancistrotellus bellus]K03
    |    |--P. difficilis (Mello-Leitão 1931) [=*Batomites difficilis]K03
    |    |--P. granulosissimus Mello-Leitão 1932 [incl. P. decoratus Mello-Leitão 1932]K03
    |    |--P. hatschbachi Soares 1945K03
    |    |--P. hauseri (Šilhavý 1979) [=Ancistrotellus hauseri]K03
    |    |--P. margaritatus Roewer 1931K03
    |    |--P. mendax Soares 1945K03
    |    |--P. ornatus (Mello-Leitão 1922) (see below for synonymy)K03
    |    `--P. viridigranulatus (Soares & Soares 1946)DG09 [=Ancistrotellus viridigranulatusK03]
    `--Neoancistrotus Mello-Leitão 1927 [incl. Neomitobatoides Mello-Leitão 1933]K03
         |--*N. thiacanthus Mello-Leitão 1927 [=Ancistrotellus thiacanthus]K03
         |--N. bifurcatus (Koch 1839)DG09 (see below for synonymy)
         |--N. bipustulatus (Mello-Leitão 1940) [=Ancistrotellus bipustulatus]K03
         |--N. bristowei (Mello-Leitão 1924) [=Ancistrotus bristowei, Ancistrotellus bristowei]K03
         |--N. dubius (Mello-Leitão 1932) [=Ancistrotellus dubius]K03
         |--N. gracilis (Roewer 1917) [=Ancistrotus gracilis, Ancistrotellus gracilis]K03
         |--N. guapimirim (Soares & Soares 1946) [=Ancistrotellus guapimirim]K03
         |--N. intermedius (Mello-Leitão 1936) [=Ancistrotellus intermedius]K03
         |--N. maculipalpi (Soares & Soares 1948) (see below for synonymy)K03
         |--N. nigroides (Soares & Soares 1947) [=Ancistrotellus nigroides]K03
         |--N. obscurus (Mello-Leitão 1931) (see below for synonymy)K03
         `--N. rosai (Mello-Leitão 1933) [=*Neomitobatoides rosae, Ancistrotellus rosae, Neoancistrotus rosae]K03

*Despirus parvulus (Roewer 1927) [=Discocyrtus parvulus, Discocyrtoides parvulus; incl. Discocyrtoides granulosus Mello-Leitão 1936, *Ibarroides minutus Mello-Leitão 1932, Discocyrtoides minutus, *Segadasius monticola Mello-Leitão 1949]K03

Dyscocyrtoides nigricans (Mello-Leitão 1922) [=Ancistrotus nigricans, *Mitobatoides nigrcans, M. nigrigans (l. c.); incl. Despirus piracicabensis Piza 1938, De. ustus Mello-Leitão 1937, *Discocyrtoides violaceus Mello-Leitão 1923, Di. ypirangae Mello-Leitão 1923]K03

Metamitobates Roewer 1913 [incl. Ancistrotellus Roewer 1923, Ancistrotus Koch 1839 non Audinet-Serville 1832, Ancystrotus (l. c.), Neobourguyia Mello-Leitão 1940, Neomitobates Roewer 1913]K03

Metamitobates squalidus (Perty 1833)MKM22 [=Goniosoma squalidumK03, *Ancistrotellus squalidusK03, *Ancistrotus squalidusK03; incl. *Metamitobates genusulphureus Roewer 1913K03, Metamit. genusulfureas (l. c.)K03, Metamit. genusulfureus (l. c.)K03, *Neomitobates gracilipes Roewer 1913K03, Metamit. gracilipesK03, *Neobourguyia rosai Mello-Leitão 1940K03, Despiroides rosaiK03, Discocyrtoides rosaiK03, Ancistrotus urceolaris Bertkau 1880K03, Neomitobates venustus Mello-Leitão 1933K03, Metamibotates (l. c.) venustusK03]

Mitobates Sundevall 1833K03 [incl. Despiroides Mello-Leitão 1932K03, Dolichoscelis Hope 1836K03, KA-Z11, Metaroeweria Mello-Leitão 1936K03]

*Mitobates triangulus Sundevall 1833K03 [incl. Goniosoma conspersum Perty 1833K03, Ancistrotus conspersusK03, Mitobates conspersusDG09, *Dolichoscelis haworthii Hope 1837K03, *Metaroeweria insignis Mello-Leitão 1936K03, Neoancistrotus nigromaculatus Mello-Leitão 1931K03, Ancistrotellus nigromaculatusK03, Mitobates stygnoides Mello-Leitão 1923K03, *Despiroides xanthophthalmus Mello-Leitão 1932K03, Discocyrtoides xanthophthalmusK03]

Neoancistrotus bifurcatus (Koch 1839)DG09 [=Ancistrotus bifurcatusK03, Ancistrotellus bifurcatusK03, Dolichoscelis bifurcatusK03, Goniosoma bifurcatumK03; incl. Ancistrotellus anomalus Mello-Leitão 1935K03, Neoancistrotus anomalusK03, Prosancistrotus communis Mello-Leitão in Soares 1945 (n. n.)K03, Ancistrotellus perlatus Mello-Leitão 1932K03, N. perlatusK03, Ancistrotellus tijucae Soares 1944K03, N. tijucaeK03]

Neoancistrotus maculipalpi (Soares & Soares 1948) [=Ancistrotellus maculipalpi; incl. A. gracilioides Mello-Leitão in Soares 1945 (n. n.), A. luteimanu Mello-Leitão in Soares 1945 (n. n.)]K03

Neoancistrotus obscurus (Mello-Leitão 1931) [=Ancistrotellus obscurus; incl. A. melacanthus Mello-Leitão 1932, Neoancistrotus melacanthus]K03

Promitobates Roewer 1913K03 [incl. Batomites Mello-Leitão 1931 (nom. inv.)K03, KA-Z11, Leonardosia Mello-Leitão 1935K03, Promitobatoides Mello-Leitão 1927K03]

Promitobates ornatus (Mello-Leitão 1922) [=Neomitobates ornatus, *Promitobatoides ornatus; incl. Batomites heteracanthus Mello-Leitão 1935, Promitobates heteracanthus, B. intermedius Mello-Leitão 1935, Neoancistrotus nigripes Mello-Leitão 1935, Ancistrotellus nigripes, *Leonardosia nitida Mello-Leitão 1935, Promitobates nitidus, Batomites nitidus Mello-Leitão 1940 non Leonardosia nitida Mello-Leitão 1935, B. spitzi Mello-Leitão 1931]K03

*Type species of generic name indicated

References

[DG09] DaSilva, M. B., & P. Gnaspini. 2009. A systematic revision of Goniosomatinae (Arachnida: Opiliones: Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics 23: 530–624.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[KA-Z11] Kury, A. B., & M. A. Alonso-Zarazaga. 2011. Addenda and corrigenda to the “Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)”. Zootaxa 3034: 47–68.

[MKM22] Medrano, M., A. B. Kury & A. C. Mendes. 2022. Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae). Zoological Journal of the Linnean Society 195 (2): 585–672.

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