Belongs within: Muridae.
Contains: Maxomys, Niviventer, Rattus, Apomys, Hydromyini, Conilurini, Otomys, Aethomys, Lemniscomys, Apodemus, Mus, Praomys.
The Murinae are a major clade of Old World rats and mice supported by molecular data and features of the dentition.
<==Murinae [Hydromyinae, Pseudomyinae]L07 |--+--Micromys Dehne 1841FS15, P04 | | `--M. minutus (Pallas 1771)KJ08 | `--+--MaxomysFS15 | `--+--+--HadromysFS15 | | | |--H. humeiFS15 | | | `--H. yunnanensisFS15 | | `--+--+--NiviventerFS15 | | | `--+--Tonkinomys daovantieniFS15 | | | `--Chiromyscus chiropusFS15 | | `--+--+--Anonymomys mindorensisFS15 | | | `--Dacnomys millardiFS15 | | `--LeopoldamysFS15 | | |--L. neilliFS15 [=Rattus neilliBP87] | | `--+--L. sabanus (Thomas 1887)FS15, JK05 | | `--+--L. siporanusFS15 | | `--+--L. milletiFS15 | | `--+--L. ciliatusFS15 | | `--L. edwardsiFS15 | `--+--+--Lenothrix canusFS15 [=Rattus canusBP87] | | |--+--Eropeplus canusFS15, M82 | | | `--Lenomys meyeriFS15 | | |--+--Pithecheirops otionFS15 | | | `--PithecheirFS15 | | | |--P. melanurusFS15 | | | `--P. parvusFS15 | | `--+--+--Melasmothrix naso Miller & Hollister 1921FS15, M82 | | | `--TateomysFS15 | | | |--T. macrocercus Musser 1982M82 | | | `--T. rhinogradoidesM82 | | `--+--EchiothrixFS15 | | | |--E. centrosaFS15 | | | `--E. leucuraM82 | | `--MargaretamysFS15 | | |--M. parvusFS15 | | `--+--M. beccariiFS15 | | `--M. elegansFS15 | `--+--+--Srilankamys ohiensisFS15 | | `--Saxatilomys paulinaeFS15 | `--+--BerylmysFS15 | | |--B. berdmoreiFS15 | | |--B. bowersi (Anderson 1879)K92 | | |--B. mackenzieiFS15 | | `--B. manipulusFS15 | `--+--+--+--Nesokia Gray 1842FS15, G91 | | | | |--*N. indica (Gray & Hardwicke 1830) (see below for synonymy)G91 | | | | `--N. bunniiFS15 | | | `--+--Kadarsanomys sodyiFS15 [=Lenothrix canus sodyiBP87] | | | `--Bandicota Gray 1873FS15, G91 | | | | i. s.: B. malabaricaJK05 | | | |--B. bengalensis (Gray & Harwicke 1833)FS15, K92 | | | `--+--B. indica (Bechstein 1800)FS15, G91 [=Mus indicusG91, Nesokia indicaG91] | | | `--B. savileiFS15 | | `--+--RattusFS15 | | `--+--Diplothrix legatus (Thomas 1906)FS15, I92 | | `--TaeromysFS15 | | |--+--T. callitrichusFS15 | | | `--T. microbullatusFS15 | | `--+--T. arcuatusFS15 | | |--T. celebensisFS15 | | |--T. hamatusFS15 | | |--T. punicansFS15 | | `--T. taeraeFS15 | `--+--BullimusFS15 | | | i. s.: B. barkeriM82 | | | B. raboriM82 | | |--B. luzonicusFS15 | | `--+--B. bagobusFS15 | | `--B. gamayFS15 | `--+--SundamysFS15 | | |--S. infraluteusFS15 | | `--+--S. maxiFS15 | | `--S. muelleriFS15 | `--+--BunomysFS15 | | |--B. andrewsiM82 | | |--B. chrysocomusM82 | | |--B. coelestisFS15 | | |--B. fratrorumFS15 | | |--B. heinrichiIT07 | | |--B. penitus (Miller & Hollister 1921) [=Rattus penitus]K92 | | `--B. prolatusFS15 | `--+--Palawanomys furvusFS15 | `--+--ParuromysFS15 | | |--P. dominatorM82 | | `--P. ursinusIT07 | `--PapagomysFS15 | |--P. theodorverhoeveniFS15 | `--+--P. armandvilleiFS15 | `--Komodomys rintjanusFS15 `--+--+--+--HaeromysFS15 | | | |--H. pusillusFS15 | | | `--+--H. margarettaeFS15 | | | `--H. minahassaeFS15 | | `--+--HapalomysFS15 | | | |--H. delacouriFS15 | | | `--H. longicaudatusFS15 | | `--ChiropodomysFS15 | | |--C. calamianensisFS15 | | |--C. gliroidesFS15 | | |--C. karlkoopmaniFS15 | | |--C. majorFS15 | | |--C. muroidesFS15 | | `--C. pusillusFS15 | `--+--+--ApomysFS15 | | `--+--Archboldomys Musser 1982FS15, M82 | | | |--A. musseriFS15 | | | `--+--*A. luzonensis Musser 1982M82, FS15, M82 | | | `--A. kalingaFS15 | | `--+--RhynchomysFS15 | | | |--R. banahaoFS15 | | | |--R. isarogensisM82 | | | |--R. soricoidesM82 | | | `--R. tapulaoFS15 | | `--ChrotomysFS15 | | |--C. silaceusFS15 | | `--+--C. sibuyanensisFS15 | | `--+--C. gonzalesiFS15 | | `--+--C. mindorensisFS15 | | `--C. whiteheadiFS15 | `--+--+--+--HydromyiniFS15 | | | `--ConiluriniFS15 | | `--+--MammelomysFS15 | | | |--M. lanosus (Thomas 1922)APB99 | | | `--M. rattoides (Thomas 1922)APB99 | | `--+--MallomysFS15 | | | |--M. aroaensis (De Vis 1907)APB99 | | | |--M. gunungFS15 | | | |--M. istapantapFS15 | | | `--M. rothschildiFS15 | | `--+--Abeomelomys sevia (Tate & Archbold 1935)FS15, APB99 | | `--PogonomelomysFS15 | | | i. s.: P. ruemmleriL73 | | | P. seviaIT07 | | |--P. mayeri (Rothschild & Dollman 1932)FS15, APB99 | | `--+--P. brassiFS15 [=P. bruijni brassiAPB99] | | `--P. bruijnii (Peters & Doria 1876)FS15, APB99 | `--+--+--LorentzimysFS15 | | | |--L. alticolaL73 | | | `--L. nouhuysiFS15 | | `--+--Anisomys imitatorFS15 | | `--Xenuromys barbatus (Milne-Edwards 1900)FS15, APB99 | `--+--+--MacruromysFS15 | | | |--M. elegansFS15 | | | `--M. majorFS15 | | `--ChiruromysFS15 | | |--C. forbesiFS15 | | `--+--C. lamiaFS15 | | `--C. vatesFS15 | `--+--+--HyomysFS15 | | | |--H. dammermaniFS15 | | | `--H. goliathFS15 | | `--CoryphomysFS15 | | |--C. buehleri Schaub 1937KAB91 | | `--C. musseriFS15 | `--+--Spelaeomys florensisFS15 | `--Pogonomys Milne Edwards 1877FS15, WA81 | | i. s.: P. mollipilosus Peters & Doria 1881WA81 | |--+--P. macrourus (Milne-Edwards 1877)FS15, APB99 | | `--P. sylvestris Thomas 1920FS15, APB99 | `--+--P. championiFS15 | `--+--P. fergussoniensisFS15 | `--P. loriae Thomas 1897FS15, APB99 `--+--+--+--Diomys crumpiFS15 | | `--+--CremnomysFS15 | | | |--C. cutchicusFS15 | | | `--+--C. blanfordiIT07 [=Madromys blanfordiFS15] | | | `--C. elviraFS15 | | `--MillardiaFS15 | | |--M. gleadowiFS15 | | |--M. kathleenaeFS15 | | |--M. kondanaFS15 | | `--M. meltadaFS15 | `--+--OtomysFS15 | `--+--Golunda elliotiFS15 | `--+--+--Lamottemys okuensisFS15 | | `--+--OenomysFS15 | | | |--O. hypoxanthusFS15 | | | `--O. ornatusFS15 | | `--ThamnomysFS15 | | |--T. venustusFS15 | | `--+--T. kempiFS15 | | `--T. schoutedeniFS15 | `--+--+--Stochomys longicaudatusFS15 | | `--+--DephomysFS15 | | | |--D. defuaFS15 | | | `--D. eburneaIT07 | | `--HybomysFS15 | | |--H. badiusFS15 | | |--H. basiliiFS15 | | |--H. eisentrautiIT07 | | |--H. lunarisFS15 | | |--H. planifronsFS15 | | |--H. trivirgatusFS15 | | `--H. univittatusFS15 | `--+--AethomysFS15 | `--+--DasymysFS15 | | |--D. foxiFS15 | | |--D. incomtusFS15 | | |--D. montanusFS15 | | |--D. nudipesFS15 | | `--D. rufulusFS15 | `--+--DesmomysFS15 | | |--D. harringtoniFS15 | | `--D. yaldeniFS15 | `--+--+--MylomysFS15 | | | |--M. dybowskiiFS15 | | | `--M. rexFS15 | | `--Pelomys Peters 1852FS15, P04 | | |--P. campanaeFS15 | | |--P. fallax (Peters 1852)K92 | | |--P. hopkinsiFS15 | | |--P. isseliFS15 | | |--P. minorFS15 | | `--P. rexBP87 | `--+--Rhabdomys pumilioFS15 | `--+--LemniscomysFS15 | `--ArvicanthisFS15 | | i. s.: A. somalicusIT07 | |--+--A. nairobaeFS15 | | `--+--A. blickiFS15 | | `--A. neumanniFS15 | `--+--A. abyssinicusFS15 | `--+--A. rufinusFS15 | `--+--A. ansorgeiFS15 | `--A. niloticus (Desmarest 1822)FS15, K92 `--+--Leimacomys buettneriFS15 `--+--+--VandeleuriaFS15 | | | i. s.: V. dumeticolaL07 | | |--V. noltheniiFS15 | | `--+--V. nilagiricaFS15 | | `--V. olearaceaFS15 | `--Vernaya Anthony 1941FS15, L07 [=Octopodomys Sody 1941L07] | |--*V. fulva (Allen 1927) (see below for synonymy)L07 | |--V. gigantaL07 | |--V. prefulvaL07 | |--V. pristinaL07 | `--V. wushanicaL07 `--+--+--+--ApodemusFS15 | | `--TokudaiaFS15 | | |--T. muenninki (Johnson 1946)FS15, I92 [=T. osimensis muenninkiI92] | | `--+--T. osimensis (Abe 1934)FS15, I92 | | `--T. tokunoshimensisFS15 | `--+--+--Muriculus imberbisFS15 | | `--MusFS15 | `--MalacomysFS15 | | i. s.: M. lukolelaeIT07 | | M. verschureniIT07 | |--M. edwardsiFS15 | `--+--M. cansdaleiFS15 | `--M. longipesFS15 `--+--+--MyomyscusFS15 | | |--M. yemeniFS15 | | `--+--M. angolensisFS15 | | `--M. brockmaniFS15 | `--StenocephalemysFS15 | |--S. albocaudataFS15 | `--+--S. griseicaudaFS15 | `--+--S. albipesFS15 | `--S. ruppiFS15 `--+--+--+--+--‘Praomys’ delectorumFS15 | | | `--‘Praomys’ morioFS15 | | `--+--‘Mastomys’ pernanusFS15 | | `--‘Mastomys’ shortridgeiFS15 | `--+--Heimyscus fumosusFS15 | `--HylomyscusFS15 | |--+--H. baeriFS15 | | `--+--H. carillusFS15 | | `--+--H. aetaFS15 | | `--H. grandisFS15 | `--+--H. denniaeFS15 | `--+--H. stellaFS15 | `--+--H. alleniFS15 | `--H. parvusFS15 `--+--PraomysFS15 `--+--+--‘Myomyscus’ verreauxiiFS15 | `--+--ZelotomysFS15 | | |--Z. hildegardeaeFS15 | | `--Z. woosnamiFS15 | `--+--Colomys goslingiFS15 | `--Nilopegamys plumbeusFS15 `--MastomysFS15 | i. s.: M. angolensisIT07 | M. hildebrandtiiIT07 | M. verheyeniIT07 |--M. kollmannspergeriFS15 `--+--M. hubertiFS15 `--+--M. couchaFS15 `--+--M. natalensisFS15 `--+--M. awashensisFS15 `--M. erythroleucusFS15 Murinae incertae sedis: Progonomys cathalaiL78 Rhagamys Forsyth-Major 1905AC98 `--*R. orthodon (Hensel 1856)AC98 Paraethomys Petter 1968L78, P04 |--P. anomalusL78 `--P. miocaenicusL78 Epimys ravus (Robinson & Kloss 1916) [=Rattus ravus]T37 Celaenomys silaceusM82 Neohydromys fuscusL73, IT07 Mayermys ellermaniL73, IT07 KritimysL78
*Nesokia indica (Gray & Hardwicke 1830) [=Arvicola indica non Mus indicus Bechstein 1800 (not preoc. if in dif. gen), N. hardwickei (Gray 1837)]G91
*Vernaya fulva (Allen 1927) [=Chiropodomys fulvus, *Octopodomys fulvus; incl. Vernaya foramena Wang, Hu & Chen 1980]L07
*Type species of generic name indicated
References
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[APB99] Aplin, K. P., J. M. Pasveer & W. E. Boles. 1999. Late Quaternary vertebrates from the Bird’s Head Peninsula, Irian Jaya, Indonesia, including descriptions of two previously unknown marsupial species. Records of the Western Australian Museum Supplement 57: 351–387.
[BP87] Burton, J. A., & B. Pearson. 1987. Collins Guide to the Rare Mammals of the World. Collins: London.
[FS15] Faurby, S., & J.-C. Svenning. 2015. A species-level phylogeny of all extant and late Quaternary extinct mammals using a novel heuristic-hierarchical Bayesian approach. Molecular Phylogenetics and Evolution 84: 14–26.
[G91] Groves, C. P. 1991. A Theory of Human and Primate Evolution revised ed. Clarendon Press: Oxford.
[IT07] Isaac, N. J. B., S. T. Turvey, B. Collen, C. Waterman & J. E. M. Baillie. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PloS One 2 (3): e296.
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[KAB91] Kitchener, D. J., K. P. Aplin & Boeadi. 1991. A new species of Rattus from Gunung Mutis, south west Timor Island, Indonesia. Records of the Western Australian Museum 15 (2): 445–461.
[K92] Klompen, J. S. H. 1992. Phylogenetic relationships in the mite family Sarcoptidae (Acari: Astigmata). Miscellaneous Publications, Museum of Zoology, University of Michigan 180: i–vi, 1–154.
[KJ08] Krämer, M., & M. Jentzsch. 2008. Kleinsäuger-Vorkommen aus dem Raum Zeitz—eine vergleichende Studie. Mauritiana 20 (2): 411–427.
[L78] Lavocat, L. 1978. Rodentia and Lagomorpha. In: Maglio, V. J., & H. B. S. Cooke (eds) Evolution of African Mammals pp. 69–89. Harvard University Press: Cambridge (Massachusetts).
[L73] Lidicker, W. Z., Jr. 1973. A phenetic analysis of some New Guinea rodents. Systematic Zoology 22 (1): 36–45.
[L07] Lunde, D. P. 2007. Vernaya fulva. Mammalian Species 806: 1–3.
[M82] Musser, G. G. 1982. Results of the Archbold Expeditions. No. 110. Crunomys and the small-bodied shrew rats native to the Philippine Islands and Sulawesi (Celebes). Bulletin of the American Museum of Natural History 174 (1): 1–95.
[P04] Popov, V. V. 2004. Pliocene small mammals (Mammalia, Lipotyphla, Chiroptera, Lagomorpha, Rodentia) from Muselievo (north Bulgaria). Geodiversitas 26 (3): 403–491.
[T37] Troughton, E. Le G. 1937. The status of “Mus” novaehollandiae Waterhouse, and allied forms. Records of the Australian Museum 20: 185–190.
[WA81] Watts, C. H. S., & H. J. Aslin. 1981. The Rodents of Australia. Angus & Robertson Publishers.