Male Myospila meditabunda, copyright James K. Lindsey.

Belongs within: Muscidae.
Contains: Brontaeini, Eginiini, Helina, Mydaea, Phaonia.

The Mydaeinae are a group of muscid flies with a bare anepimeron, long anterior presutural dorsocentral bristles, and rounded lower calypters. These features are shared with the subfamily Phaoniinae and the phylogenetic analysis of calyptrate flies by Kutty et al. (2010) suggests that the two subfamilies are not reciprocally monophyletic. Members of the combined group of the two subfamilies include the cosmopolitan genus Myospila, characterised by a bare prosternum and setulae on the dorsal surface of vein R4+5 (Couri 2010).

<==Mydaeinae [Phaoniinae, Phaoniini]KP10
| `--+--MydaeaKP10
| `--+--PhaoniaKP10
| `--Hebecnema Schnabl 1889KP10, F92
| |--*H. umbratica (Meigen 1826) [=Anthomyia umbratica, Aricia (*Hebecnema) umbratica]C10
| |--H. affinis Malloch 1921F92
| |--H. alba Hsue 1983F92
| |--H. fulva (Bigot 1885) [=Spilogaster fulvus]H72
| `--H. fumosa (Meigen 1826) [=Anthomyia fumosa]F92
`--Myospila Rondani 1856KP10, F92 (see below for synonymy)
|--*M. meditabunda (Fabricius 1782)C10 (see below for synonymy)
| |--M. m. meditabundaF92
| |--M. m. angustifrons Malloch 1922F92
| `--M. m. brunettiana Enderlein 1927F92
|--M. argentata (Walker 1857) (see below for synonymy)F92
|--M. bina (Wiedemann 1830) [=Anthomyia bina]F92
|--M. effeminataF92
|--M. femorata (Malloch 1935) [=Xenosina femorata]F92
|--M. flavibasis (Malloch 1925) [=Xenosia flavibasis]F92
|--M. fuscicoxa (Li 1980) [=Eumyiospila fuscicoxa]F92
|--M. laevis (Stein 1900) (see below for synonymy)F92
|--M. lasiophthalma (Emden 1965) [=Xenosina lasiophthalma]F92
|--M. lauta (Stein 1918) [=Mydaea lauta]F92
|--M. lenticeps (Thomson 1869) [=Anthomyia lenticeps]F92
`--M. tenax (Stein 1918) [=Mydaea tenax]F92

Mydaeinae incertae sedis:
Opsolasia Coquillett 1910F92
`--O. orichalceaHV87
Lasiopelta Malloch 1928 [incl. Pandleburyia Malloch 1928]F92
|--*L. orientalis Malloch 1928F92
`--L. longicornis (Stein 1915) [=Mydaea longicornis]F92
Dichaetomyia Malloch 1921 [Dichaetomyiini]F92
|--D. emdeni Pont 1969 [incl. *D. polita Malloch 1921 (preoc.)]C10
|--D. alterna (Stein 1915) [=Mydaea alterna]F92
|--D. antennata (Stein 1918) [=Mydaea antennata]F92
|--D. apicalis (Stein 1904) [=Spilogaster apicalis]F92
|--D. aureomarginata Emden 1965 [=D. nubiana aureomarginata]F92
|--D. bibax (Wiedemann 1830) [=Anthomyia bibax]F92
|--D. femorata (Stein 1915) [=Mydaea femorata]F92
|--D. flavipalpis (Stein 1915) [=Mydaea flavipalpis]F92
|--D. flavocaudata Malloch 1925F92
|--D. keiseri Emden 1965F92
|--D. luteiventris (Rondani 1873) (see below for synonymy)F92
|--D. monticola Emden 1965F92
|--D. pallicornis (Stein 1915) [=Mydaea pallicornis]F92
|--D. pallitarsis (Stein 1909) [=Mydaea pallitarsis]F92
|--D. pectinipesF92
|--D. quadrata (Wiedemann 1824) [=Anthomyia quadrata]F92
`--D. setifemur Malloch 1928F92
Metopomyia Malloch 1922 [Metopomyiini]C10
`--*M. atropunctipes Malloch 1922C10
Beccimyia Pont 1969C10
`--*B. papuana Pont 1969C10
Buccophaonia Emden 1965C10
`--*B. subcostalis Emden 1965C10
Hennigiola Pont 1969C10
`--*H. setulifera (Stein 1906) [=Spilogaster setulifera]C10
Idiohelina Malloch 1921C10
`--*I. nubeculosa Malloch 1921C10
Phaonina Emden 1965C10
`--*P. corbetti (Malloch 1931) [=Phaonia corbetti]C10
Prohardyia Pont 1969 [=Hardyia Malloch 1926 (preoc.)]C10
`--*P. carinata (Stein 1910) [=Mydaea carinata, *Hardyia carinata]C10
Chaetopapuaia Vockeroth 1972C10
`--*C. setifrons Vockeroth 1972C10
Graphomya Robineau-Desvoidy 1830C10
|--*G. maculata (Scopoli 1763)F92 [=Musca maculataF92, Curtonevra maculataL49]
| |--G. m. maculataF92
| `--G. m. tienmushanensis Ôuchi 1939F92
|--G. idessaSV10
|--G. mellina Stein 1909F92
|--G. paucimaculata Ôuchi 1938 [=Graphomyia paucimaculata]F92
`--G. rufitibia Stein 1918 [=Graphomyia rufitibia]F92
Gymnopapuaia Vockeroth 1972C10
`--*G. albicornis (Walker 1864) [=Aricia albicornis]C10
Helinomydaea Vockeroth 1972C10
|--*H. acrostichalis Vockeroth 1972C10
`--H. punctiventrisC10
Papuaia Malloch 1921C10
|--P. curvinervis (Stein 1900) (see below for synonymy)C10
`--P. asternataC10
Papuaiella Vockeroth 1972C10
`--*P. ponti Vockeroth 1972C10

Dichaetomyia luteiventris (Rondani 1873) [=Anthomyia luteiventris; incl. D. lineata (Stein 1904), D. nubiana (Bigot 1885)]F92

Myospila Rondani 1856KP10, F92 [incl. Eumyiospila Malloch 1926F92, Helinella Malloch 1926F92, Parapictia Pont 1968F92, Sinomusina Séguy 1937F92, Xenosia Malloch 1921F92, Xenosina Malloch 1925F92]

Myospila argentata (Walker 1857) [=Aricia argentata; incl. Eumydaea attenta Stein 1918, Spilogaster pellucida Stein 1928, Eumyiospila spinifemarata Malloch 1926]F92

Myospila laevis (Stein 1900) [=Spilogaster laevis; incl. S. arminervis Stein 1900, Xenosia scutellaris Malloch 1935, X. tarsalia Malloch 1935]F92

*Myospila meditabunda (Fabricius 1782)C10 [=Musca meditabundaF92; incl. Curtonevra anthomydea Bigot 1887H72, C. nigriceps Bigot 1887H72]

Papuaia curvinervis (Stein 1900) [=Spilogaster curvinervis; incl. S. rufescens Stein 1900, *Papuaia rufescens]C10

*Type species of generic name indicated


[C10] Couri, M. S. 2010. Key to the Australasian and Oceanian genera of Muscidae (Diptera). Revista Brasileira de Entomologia 54 (4): 529–544.

[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.

[H72] Huckett, H. C. 1972. Notes on Bigot’s North American type-specimens at the University Museum, Oxford (Diptera: Anthomyiidae, Muscidae). Pan-Pacific Entomologist 48 (2): 81–85.

[HV87] Huckett, H. C., & J. R. Vockeroth. 1987. Muscidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1115–1131. Research Branch, Agriculture Canada.

[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine’s fly. Systematic Entomology 35: 614–635.

[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.

[SV10] Savage, J., & J. R. Vockeroth. 2010. Muscidae (house flies, stable flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1281–1295. NRC Research Press: Ottawa.

Leave a comment

Your email address will not be published. Required fields are marked *