Inoperna plicata, from here.

Belongs within: Pteriomorphia.
Contains: BrachidontesMusculus, Modiolaria, Modiolus, Perna, Lithophaga, Mytilus, Crenella.

The Mytilidae are the mussels, a family of originally epifaunal, byssally-attached bivalves. Identifiable examples of this family are known in the fossil record from the Devonian (Skelton & Benton 1993). Free-living species are commonly nestling, forming distinct colonies. The beak of the shell is at the anterior end in Mytilinae but slightly behind the anterior in Modiolinae. Members of the subfamily Lithophaginae have become borers into their rock substrate. The Pacific genus Adula is characterised by an elongate, cylindrical shell with the beak behind the anterior margin and a thickened anterodorsal margin. The Crenellinae include byssally attached, infaunal and boring species in which the lobes of the mantle are anteroventrally fused (Beesley et al. 1998).

    |  |    |--ModiolusPC11
    |  |    |--Modiolula phaseolinaPC11
    |  |    |--PernaH00
    |  |    |--AdipicolaBRW98
    |  |    |--Lioberus castaneus (Say 1822)BRW98, S-S90
    |  |    `--ModiolusiaBRW98
    |  `--LithophaginaeCS04
    |       |--LithophagaPC11
    |       |--AmygdalumF26
    |       |    |--A. beddomei Iredale 1924MG-H11
    |       |    `--A. watsoni Smith 1895MG-H11
    |       |--Zelithophaga Finlay 1927F26
    |       |    |--*Z. truncata (Gray 1843)P61 [=Lithodomus truncatusP61, Lithophaga truncataF26]
    |       |    `--Z. nelsoniana [=Lithophaga nelsoniana]F26
    |       |--AdulaH00
    |       |    |--A. cinnamomea [=Lithodomus cinnamomeus, L. cinnamominus]C64
    |       |    |--A. falcata [incl. A. gruneri]C64
    |       |    `--A. stylina Carpenter 1864C64
    |       `--Inoperna Conrad in Kerr 1875 [incl. Pharomytilus Rollier 1914-1915]H00
    |            |--*I. carolinensis Conrad in Kerr 1875 [=Modiolus (Inoperna) carolinensis]H00
    |            |--I. lilliputensis Hodges 2000H00
    |            `--I. plicata Sowerby 1819 (see below for synonymy)H00
       |    |--M. lineatusCS77
       |    `--M. monterosatoiP75
          |    |--MytilusPC11
          |    |--Limnoperna fortuneiG88, M96
          |    |--AustromytilusBRW98
          |    |    |--A. australisBRW98
          |    |    `--A. rostratusBRW98
          |    `--XenostrobusBRW98
          |         |--X. atrata (Lischke 1871)XZ10
          |         |--X. pulexHS01
          |         `--X. securisHS01
               |    |--M. glaberrima (Dunker 1857)S-S90
               |    `--M. senhousia (Benson in Cantor 1842)PC11, HJ08
               `--Gregariella Monterosato 1884P61 [incl. Trichomusculus Iredale 1924P61, F26]
                    |--*G. sulcata [=Modiolus sulcatus]P61
                    |--G. barbata (Reeve 1858)P61 [=Lithodomus barbatusH09, Modiolaria barbataF26, *Trichomusculus barbatusF26]
                    `--G. petagnaeBBB-S95
Mytilidae incertae sedis:
  Geukensia demissaGW02
    |--G. d. demissaDK89
    `--G. d. granosissima Sowerby 1914DK89
  Aulacomya Moerch 1853P61
    |--*A. magellanica (Lamarck 1819)P61, F26 [=Mytilus magellanicusP61]
    |--A. ater (Molina 1782)GO78
    `--A. maoriana (Iredale 1915)P61 [=Mytilus (Aulacomya) maorianusF26, Brachyodontes maorianusF26]
  Trichomya Ihering 1924F26
    |--T. hirsuta (Lamarck 1819)F26, H09 [=Mytilus hirsutusF26, Brachyodontes hirsutusH09]
    |--T. huttoni [=Mytilus huttoni]F26
    |--T. rostrata [=Mytilus rostratus]F26
    `--T. torquata (Marshall 1918) [=Mytilus torquatus]F26
  Crenomytilus grayanus (Dunker 1853)VKK10
  Solamen Iredale 1924DK08
    |  i. s.: S. spectabilis (Adams 1862)XZ10
    `--S. (Exosiperna Iredale 1929)DK08
         `--S. globularis (Tate 1886) [=Crenella globularis]DK08
    |--B. fusca (Gmelin 1791)BW09
    |--B. silicula (Lamarck 1819)BW09
    `--B. vagina (Lamarck 1819)K90
  Septifer Recluz 1848P61
    |--*S. bilocularis (Linnaeus 1758)P61 (see below for synonymy)
    |--S. bifurcatus [=Mytilus bifurcatus]C64
    |--S. cumingianus [=Mytilus cumingianus]C64
    |--S. curvirostrisG31
    `--‘Mytilus’ mutabilis Gould 1861C64
  Stavelia Gray 1858W60
    `--S. horrida Récluz 1852WMJ09 [=Mytilus horridusH09]
  Falcimytilus Cox 1937W60
  Ryenella Fleming 1959P61
    |--*R. impacta (Hermann 1782) [=Mytilus impactus]P61
    `--R. cumingiana (Dunker 1857) [=Musculus cumingianus]S-S90
  Dacrydium [Dacrydiinae]P61
    |  i. s.: D. vitreumHG98
    |         D. viviparumBRW98
    `--D. (Quendreda Iredale 1936)P61
         |--D. (*Q.) fabaleP61
         `--D. (Q.) pelseneeri Hedley 1906P61
  Mytilops precedens (Hall 1870)SB93
  Bathymodiolus Kenk & Wilson 1985AJH07 [BathymodiolinaeBRW98]
    `--B. breviorR01
    |--*M. plicatus (Gmelin 1791) [=Mytilus plicatus]S-S90
    `--M. hanleyi (Dunker 1882)K90
  Volsellina Newell 1942DX84
    `--V. zhesiensis Liang in Ding, Xia et al. 1984DX84

Inoperna plicata Sowerby 1819 [=Mytilus plicatus non Gmelin 1791 (not preoc. if in dif. gen.), *Pharomytilus plicatus]H00

*Septifer bilocularis (Linnaeus 1758)P61 [=Mytilus bilocularisP61, Brachyodontes (Septifer) bilocularisS-S90; incl. M. efflugensBRW98]

*Type species of generic name indicated


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