Neocomitidae

Subthurmannia boissieri, from Parent et al. (2015).

Belongs within: Perisphinctoidea.
Contains: Berriasella, Neocomites.

The Neocomitidae are a diverse assemblage of ammonites known from the Upper Tithonian of the Upper Jurassic to the Upper Hauterivian or Lower Barremian of the Lower Cretaceous. Representatives tend to become compressed and flat-sided with a tabulate or grooved venter (Wright et al. 1996).

<==Neocomitidae [Berriasellidae, Palaehoplitidae]EH19
    |--Berriasellinae [Pseudosubplanitinae]EH19
    |    |--BerriasellaWCH96
    |    |--Andiceras Krantz 1928WCH96
    |    |    `--*A. trigonostomum Krantz 1928WCH96
    |    |--Parandiceras Spath 1939WCH96
    |    |    `--*P. rota Spath 1939WCH96
    |    |--Chigaroceras Howarth 1992WCH96
    |    |    `--*C. banikense Howarth 1992WCH96
    |    |--Banikoceras Howarth 1992WCH96
    |    |    `--*B. involutum Howarth 1992WCH96
    |    |--Fierrites Cantú Chapa 1993EH19
    |    |    `--*F. alamitosensis (Aguilera in Castillo & Aguilera 1895) [=Ammonites alamitosensis]EH19
    |    |--Hegaratia Patrulius & Avram 1976EH19
    |    |    `--*H. busnardoi (Patrulius & Avram 1976) [=Corongoceras (*Hegaratia) busnardoi]EH19
    |    |--Substeueroceras Spath 1923WCH96
    |    |    |--*S. koeneni (Steuer 1897) [=Odontoceras koeneni]WCH96
    |    |    `--S. stantoniSY85
    |    |--Riasanites Spath 1923 [incl. Tauricoceras Kvantaliani & Lysenko 1979]WCH96
    |    |    |--*R. rjasanensis (Lahusen 1883) [=Ammonites rjasanensis]WCH96
    |    |    `--*Tauricoceras’ crassicostatum Kvantaliani & Lysenko 1979WCH96
    |    |--Blanfordiceras Cossmann 1907 [=Blanfordia Uhlig 1905 non Adams 1863; incl. Pseudoblanfordia Spath 1925]WCH96
    |    |    |--*B. wallichi (Gray 1832) [=Ammonites wallichi, *Blanfordia wallichi]WCH96
    |    |    `--‘Hoplites’ australis Burckhardt 1903 [=*Pseudoblanfordia australis]WCH96
    |    |--Lytohoplites Spath 1925WCH96
    |    |    |--*L. burckhardti (Mayer-Eymar in Burckhardt 1900) [=Hoplites burckhardti]WCH96
    |    |    `--L. besaireiWCH96
    |    |--Pseudosubplanites Le Hégarat 1971EH19
    |    |    |--*P. berriasensis Le Hégarat 1971EH19
    |    |    |--P. euxinusEH19
    |    |    `--P. grandisR81
    |    |--Neocosmoceras Blanchet 1922 (see below for synonymy)WCH96
    |    |    |--*N. sayni (Simionescu 1899) [=Hoplites sayni]WCH96
    |    |    |--‘Ammonites’ euthymi Pictet 1867 [=*Euthymiceras euthymi]WCH96
    |    |    |--‘Ammonites’ octagonus Blanford in Salter & Blanford 1865 [=*Octagoniceras octagonum]WCH96
    |    |    `--‘Protacanthodiscus’ transcaspius Luppov, Bodylevskii & Glazunova 1949 [=*Transcapiites transcaspius]WCH96
    |    `--Malbosiceras Grigorieva 1938WCH96 (see below for synonymy)
    |         |--*M. malbosi (Pictet 1867) [=Ammonites malbosi]WCH96
    |         |--‘Perisphinctes’ andrussovi Retowski 1893 [=*Retowskiceras andrussovi]WCH96
    |         |--‘Ammonites’ breveti Pomel 1889 [=*Pomeliceras breveti]WCH96
    |         |--‘Berriasella’ broussi Mazenot 1939 [=*Mazenoticeras broussi]WCH96
    |         |--‘Ammonites’ chaperi Pictet 1868EH19 [=*Chapericeras chaperiWCH96, *Lopeziceras chaperiEH19]
    |         `--M. paramimounumR81
    |--Endemoceratinae [Endemoceratidae, Leopoldiinae]WCH96
    |    |--Chamalocia Thieuloy 1971WCH96
    |    |    `--*C. subaenigmatica (Sayn 1907) [=Leopoldia subaenigmatica]WCH96
    |    |--Pseudofavrella Leanza & Leanza 1973WCH96
    |    |    `--*P. angulatiformis (Behrendsen 1892) [=Hoplites angulatiformis]WCH96
    |    |--Cruasiceras Busnardo 1970WCH96
    |    |    `--*C. cruasense (Torcapel 1884) [=Ammonites cruasensis]WCH96
    |    |--Acanthodiscus Uhlig 1905WCH96
    |    |    `--*A. radiatus (Bruguière 1789) [=Ammonites radiatus]WCH96
    |    |--Suboosterella Spath 1924WCH96
    |    |    `--*S. heliaca (Orbigny 1840) [=Ammonites heliacus]WCH96
    |    |--Breistrofferella Thieuloy 1971WCH96
    |    |    `--*B. castellanensis (Orbigny 1840) [=Ammonites castellanensis]WCH96
    |    |--Hannaites Imlay 1957WCH96
    |    |    `--*H. riddlensis (Anderson 1938) [=Neocomites riddlensis]WCH96
    |    |--Saynella Kilian 1910WCH96
    |    |    `--*S. clypeiformis (Orbigny 1841) [=Ammonites clypeiformis]WCH96
    |    |--Malgesaynella Busnardo 1970WCH96
    |    |    `--*M. besairei (Collignon 1949) [=Saynella besairei]WCH96
    |    |--Rodighieroites Company 1987WCH96
    |    |    `--*R. cardulus Company 1987WCH96, MBF15
    |    |--Stoicoceras Whitehouse 1927 [incl. Dicostella Busnardo 1966]WCH96
    |    |    |--*S. teutobergense (Weerth 1884) [=Ammonites (Hoplites) teutobergensis]WCH96
    |    |    `--S. tuberculatum (Roman 1933)WCH96 (see below for synonymy)
    |    |--Hatchericeras Stanton 1901 [incl. Pseudohatchericeras Leanza 1970]WCH96
    |    |    |--*H. patagonense Stanton 1901WCH96
    |    |    `--H. argentinense Stanton 1901 [=*Pseudohatchericeras argentinense]WCH96
    |    |--Leopoldia Mayer-Eymar 1887 (see below for synonymy)WCH96
    |    |    |--*L. leopoldina (Orbigny 1840) [=Ammonites leopoldinus, *Hoplitides leopoldinus, *Solgeria leopoldina]WCH96
    |    |    `--L. renevieri Baumberger 1906 [=*Renevierites renevieri]WCH96
    |    |--Distoloceras Hyatt 1900WCH96
    |    |    |--*D. hystrix (Phillips 1829) [=Ammonites hystrix]WCH96
    |    |    `--D. pavloviWCH96
    |    |--Favrella Douvillé 1909 [incl. Patagoniceras Wetzel 1960]WCH96
    |    |    |--*F. americana (Favre 1908) [=Neocomites americanus]WCH96
    |    |    |--‘Neocomites’ steinmanni Favre 1908 [=*Patagoniceras steinmanni]WCH96
    |    |    `--F. wilckensiRC02
    |    |--Neohoploceras Spath 1939 (see below for synonymy)WCH96
    |    |    |--*N. submartini (Mallada 1887) [=Ammonites submartini]WCH96
    |    |    |--N. arnoldi (Pictet & Campiche 1860)MBF15, WCH96 [=Ammonites arnoldiWCH96, *Arnoldia arnoldiWCH96]
    |    |    |--N. deperetiMBF15
    |    |    `--N. schardtiMBF15
    |    |--Karakaschiceras Thieuloy 1971WCH96
    |    |    |--*K. biassalense (Karakasch 1889) [=Hoplites biassalensis]WCH96
    |    |    |--K. attenuatum (Behrensen 1892)MBF15
    |    |    |--K. inostranzewiMBF15
    |    |    |--K. neumayri (Behrensen 1892)MBF15
    |    |    |--K. pronecostatum Felix 1891MBF15
    |    |    `--K. quadristrangulatum (Sayn 1907)MBF15
    |    `--Lyticoceras Hyatt 1900 (see below for synonymy)WCH96
    |         |--*L. cryptoceras (Orbigny 1840) [=Ammonites cryptoceras]WCH96
    |         |--L. amblygonium (Neumayr & Uhlig 1881) [=Hoplites amblygonius, *Endemoceras amblygonium]WCH96
    |         |--L. colcanapi (Collignon 1962) [=*Besairieceras colcanapi]WCH96
    |         |--L. nodosoplicatumWCH96
    |         |--L. noricumWCH96 [=Endemoceras noricumR81]
    |         |--L. pseudoregaleR81
    |         |--L. regaleWCH96 [=Endemoceras regaleR81]
    |         `--L. stevrecensis (Breskovski 1967) [=*Eleniceras stevrecensis]WCH96
    `--NeocomitinaeWCH96
         |--NeocomitesWCH96
         |--Protothurmannia Crickmay 1932WCH96
         |    `--*P. rezanoffiana Crickmay 1932WCH96
         |--Dalmasiceras Djanélidzé 1922WCH96
         |    `--*D. dalmasi (Pictet 1867) [=Ammonites dalmasi]WCH96
         |--Frenguelliceras Leanza 1945WCH96
         |    `--*F. magister Leanza 1945WCH96
         |--Raimondiceras Spath 1924 [=Pfluckeria Lisson in Lisson & Boit 1924]WCH96
         |    `--*R. raimondii (Lisson 1907) [=Hoplites raimondii, *Pfluckeria raimondii]WCH96
         |--Acantholissonia Leanza 1972WCH96
         |    `--*A. gerthi (Weaver 1931) [=Spiticeras gerthi]WCH96
         |--Cuyaniceras Leanza 1945WCH96
         |    `--*C. transgrediens (Steuer 1897) [=Odontoceras transgrediens]WCH96
         |--Calliptychoceras Spath 1924WCH96
         |    `--*C. calliptychum (Uhlig 1910) [=Neocomites calliptychus]WCH96
         |--Odontodiscoceras Spath 1924WCH96
         |    `--*O. odontodiscum (Uhlig 1910) [=Neocomites odontodiscus]WCH96
         |--Limaites Lisson in Lisson & Boit 1924WCH96
         |    `--*L. peruanus (Lisson 1907) [=Hoplites leopoldinus var. peruanum]WCH96
         |--Criosarasinella Thieuloy 1977WCH96
         |    `--*C. furcillata Thieuloy 1977WCH96
         |--Lissonia Gerth 1925WCH96
         |    `--*L. riveroi (Lisson 1907) [=Hoplites riveroi]WCH96
         |--Pseudargentiniceras Spath 1925WCH96
         |    |--*P. abscissum (Oppel in Zittel 1868) [=Ammonites abscissus]WCH96
         |    `--P. boissieri [=Berriasella (Pseudargentinoceras) boissieri]K79
         |--Sarasinella Uhlig 1905WCH96
         |    |--*S. ambigua (Uhlig 1902) [=Hoplites ambiguus]WCH96
         |    `--S. variansWCH96
         |--Subalpinites Mazenot 1939 (see below for synonymy)WCH96
         |    |--*S. fauriensis Mazenot 1939 [=*Pomeliceras fauriense]WCH96
         |    |--‘Berriasella’ jabronensis Mazenot 1939 [=*Jabronella jabronensis]WCH96
         |    `--‘Hoplites’ paquieri Simionescu 1899 [=*Erdenella paquieri]WCH96
         |--Argentiniceras Spath 1924 [incl. Andesites Gerth 1925, Boncheviceras Nikolov 1966]WCH96
         |    |--*A. malarguense (Steuer 1897) [=Odontoceras malarguense]WCH96
         |    |--‘Berriasella’ ardescensis Mazenot 1939 [=*Boncheviceras ardescensis]WCH96
         |    |--‘Perisphinctes’ loncochensis Steuer 1897 [=*Andesites loncochensis]WCH96
         |    `--A. noduliferumR81
         |--Kilianella Uhlig 1905 [incl. Luppovella Nikolov 1966, Parakilianella Sato 1961 (n. n.)]WCH96
         |    |--*K. pexiptycha (Uhlig 1882) [=Hoplites pexiptychus]WCH96
         |    |--K. roubaudianaWCH96
         |    |--K. superba (Sayn 1907) [=Thurmannia (Kilianella) superba, *Luppovella superba]WCH96
         |    `--*Parakilianella’ umazawensis Sato 1961WCH96
         |--Thurmanniceras Cossmann 1901 [=Thurmannia Hyatt 1900 non Heer 1852, Thurmannites Kilian & Reboul 1914]WCH96
         |    |  i. s.: T. otopetaWCH96
         |    |         T. pertransiensWCH96
         |    |--*T. (Thurmanniceras) thurmanni (Pictet & Campiche 1860) (see below for synonymy)WCH96
         |    `--T. (Clavithurmannia Thieuloy in Busnardo, Thieuloy & Moullade 1979)WCH96
         |         `--T. (*C.) foraticostatum Thieuloy in Busnardo, Theuloy & Moullade 1979WCH96
         `--Subthurmannia Spath 1939 (see below for synonymy)WCH96
              |--*S. fermori Spath 1939WCH96
              |--‘Berriasella’ alpillensis Mazenot 1939 [=*Tirnovella alpillensis]WCH96
              |--S. boissieriWCH96 [=Fauriella boissieriR81]
              |--‘Ammonites’ carpathicus Zittel 1868 [=*Strambergella carpathica]WCH96
              |--S. gallica (Mazenot 1939) [=Berriasella gallica, *Fauriella gallica]WCH96
              |--S. occitanicaWCH96 [=Tirnovella occitanicaR81]
              |--‘Hoplites’ retowskyi Sarasin & Schöndelmayer 1901 [=*Pseudoneocomites retowskyi]WCH96
              `--‘Tirnovella’ subalpinaR81

Leopoldia Mayer-Eymar 1887 [=Hoplitides Koenen 1902, Solgeria Uhlig 1905; incl. Renevierites Busnardo & Thieuloy 1989]WCH96

Lyticoceras Hyatt 1900 [incl. Besairieceras Collignon 1962, Eleniceras Breskovski 1967, Endemoceras Thiermann 1964]WCH96

Malbosiceras Grigorieva 1938WCH96 [incl. Chapericeras Hoedemaker 1981WCH96, Lopeziceras Frau et al. 2016EH19, Mazenoticeras Nikolov 1966WCH96, Pomeliceras Grigorieva 1938WCH96, Retowskiceras Nikolov 1966WCH96]

Neocosmoceras Blanchet 1922 [incl. Euthymiceras Grigorieva 1938, Octagoniceras Spath 1924, Transcapiites Luppov in Bogdanova, Luppov & Mikhailova 1985]WCH96

Neohoploceras Spath 1939 [incl. Arnoldia Stolley 1937 nec Cass. 1824 (ICBN) nec Massal. 1856 (ICBN) nec Mayer-Eymar 1887]WCH96

Stoicoceras tuberculatum (Roman 1933)WCH96 [=Hoplites (Neocomites) houdardi var. tuberculataWCH96, Dicostella tuberculataR81; incl. *D. pitrei Busnardo 1966WCH96]

Subalpinites Mazenot 1939 [=Pomeliceras Hoedemaker 1981 non Grigorieva 1938; incl. Erdenella Nikolov 1979, Jabronella Nikolov 1966]WCH96

Subthurmannia Spath 1939 [=Subthurmanniceras Nikolov 1960; incl. Fauriella Nikolov 1966, Pseudoneocomites Hoedemaker 1982, Strambergella Nikolov 1966, Tirnovella Nikolov 1966]WCH96

*Thurmanniceras (Thurmanniceras) thurmanni (Pictet & Campiche 1860) [=Ammonites thurmanni, *Thurmannia thurmanni, *Thurmannites thurmanni]WCH96

*Type species of generic name indicated

References

[EH19] Énay, R., & M. K. Howarth. 2019. Part L, revised, volume 3B, chapter 7: Systematic descriptions of the Perisphinctoidea. Treatise Online 120: 1–184.

[K79] Kauffman, E. G. 1979. Cretaceous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A418–A487. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[MBF15] Mourgues, F. A., L. G. Bulot & C. Frau. 2015. The Valanginian Olcostephaninae Haug, 1910 (Ammonoidea) from the Andean Lower Cretaceous Chañarcillo Basin, northern Chile. Andean Geology 42 (2): 213–236.

[R81] Rawson, P. F. 1981. Early Cretaceous ammonite biostratigraphy and biogeography. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 499–529. Academic Press.

[RC02] Riding, J. B., & J. A. Crame. 2002. Aptian to Coniacian (Early–Late Cretaceous) palynostratigraphy of the Gustav Group, James Ross Basin, Antarctica. Cretaceous Research 23: 739–760.

[SY85] Sha J.-G. & Yuan F.-S. 1985. Buchia fauna from Dong’anzhen Formation near Dong’an of Raohe County, Heilongjiang. Acta Palaeontologica Sinica 24 (6): 651–662.

[WCH96] Wright, C. W., J. H. Calloman & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

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