Checkered nerites Nerita tessellata, copyright James St. John.

Belongs within: Neritidae.

Nerita, the nerites, is a cosmopolitan genus of marine snails with an ovate to hemispherical solid shell bearing spiral ribs (Bandel 2001). Shells of the type subgenus have a thickened outer lip thickened bearing two teeth and a flattened labial area with two large teeth at the margin (Knight et al. 1960).

Characters (from Bandel 2001): Shell ovate to hemispherical, solid, with low spire, enveloping body whorl with aperture usually bearing denticulate columellar septum and crenulated interior on outer lip; ornament of spiral ribs. Inner walls resorbed. Flat callus of inner lip extending over much of base, commonly bearing teeth, folds or pimples. Operculum calcareous, usually with hinge as a flattened projection on inner side next to columellar lip inserting into muscle, supported by basal thickening and connected to a lateral ridge; patterned muscle scar present below lateral ridge. Protoconch globose, smooth, consisting of several, strongly involute whorls.

<==Nerita Linnaeus 1758VC88 (see below for synonymy)
    |--*N. (Nerita) peloronta Linnaeus 1758VC88 [=*Neritarius pelorontaKC60, *Tenare pelorontaKC60]
    |--N. (Amphinerita Martens 1887)B01 [=Odontostoma Mörch 1852 nec Turton 1829 nec d’Orbigny 1841KC60]
    |    |--N. (*A.) umlaasiana Krauss 1848B01 [=N. (Puperita) polita umlaasianaF27, *Odontostoma umlaasianumKC60]
    |    `--N. (A.) articulataB01
    |--N. (Chingua Clark & Durham 1946)KC60
    |    `--N. (*C.) chinguensis Clark & Durham 1946KC60
    |--N. (Cymostyla von Martens 1887)VC88
    |    `--N. (*C.) undata Linnaeus 1758B01 (see below for synonymy)
    |         |--N. u. undataB01
    |         `--N. u. pseudoplicata Strausz 1960B01
    |--N. (Heminerita Martens 1887)B01
    |    |--N. (H.) japonica Dunker 1859 [incl. N. (*H.) pica Gould 1850 non Gmelin 1791]B01
    |    |--N. (H.) guamensis Quoy & Gaimard 1834B01
    |    |--N. (H.) insculpta Récluz 1841B01
    |    `--N. (H.) morio Sowerby 1833B01
    |--N. (Ilynerita von Martens 1887)VC88
    |    |--N. (*I.) planospira Anton 1839VC88 (see below for synonymy)
    |    `--N. (I.) listrota Woodring 1973VC88
    |--N. (Melanerita Martens 1887)B01
    |    |--N. (M.) atramentosa Reeve 1855B01 (see below for synonymy)
    |    |--N. (M.) melanotragus (Smith 1884)B01, WG71 [=N. (Puperita) melanostragusF27]
    |    `--N. (M.) ocellata Le Guillou 1841B01
    |--N. (Pila Klein 1758)B01
    |--N. (Ritena Gray 1858)VC88
    |    `--N. (*R.) plicata Linnaeus 1758B01 (see below for synonymy)
    `--N. (Theliostyla Mörch 1852)VC88 [=Natere Gray 1858KC60]
         |--N. (*T.) albicilla Linnaeus 1758VC88 (see below for synonymy)
         |--N. (T.) ascensionensis Gmelin 1791VC88
         |--N. (T.) exuvia Linnaeus 1758VC88
         |--N. (T.) exuvioides Trechmann 1935VC88
         |--N. (T.) fortidentata Vermeij & Collins 1988VC88
         |--N. (T.) fulgurans Gmelin 1791VC88
         |--N. (T.) funiculata Menke 1851VC88
         |--N. (T.) senegalensis Gmelin 1791B01
         |--N. (T.) tampaensis Dall 1892VC88
         `--N. (T.) tessellata Gmelin 1791VC88
Nerita incertae sedis:
  N. africana Récluz 1843R43b
  N. antillarum [incl. N. praecognita]G79
  N. antiquata Récluz 1853B01
  N. apiata Récluz 1843R43a
  N. aurantiaR43b
  N. balteata Reeve 1855W93
  N. beaniana Récluz 1843R43b
  N. bergadana Vidal 1921BK03
  N. bernhardiC64
  N. bidens Linnaeus 1758L58
  N. brasilianaR43b
  N. chamaeleon Linnaeus 1758 [incl. N. bizonalis Lamarck 1816]W93
  N. cingulata Reuss 1854BK03
  N. conoideaO75
  N. costata Gmelin 1791 [incl. N. grossa Born 1780 non Linnaeus 1758, N. scabricosta de Lessert 1841]W93
  N. cuprina Récluz 1843R43a
  N. dainellii Cossmann 1925B01
  N. donovana Récluz 1843R43a
  N. doreyana Quoy & Gaimard 1834B01
  N. dubiaR42
  N. gemmata Pethö 1906BK03
  N. gigantea Bellardi & Michelotti 1840B01
  N. grayana Récluz 1843R43b
  N. grossa Linnaeus 1758H09
  N. guillouanaF66
  N. guttataR43b
  N. hadra Woodring 1973VC88
  N. heberti (Szöts 1953) [=Neritina heberti]B01
  N. hindsii Récluz 1843R43b
  N. jayanca Olsson 1944BK03
  N. jovis Récluz 1843R43a
  N. kaffraria Woods 1906BK03
  N. lacustris Linnaeus 1758L58
  N. leachii Récluz in Guérin 1841R43b
  N. lineata Gmelin 1791WG71
  N. littoralis Linnaeus 1758L58
  N. lugubrisR43a
  N. mertoniana Récluz 1843R43a
  N. michaudiana Récluz 1841R43a
  N. nebulata Récluz 1843R43a
  N. oligopleura Dall & Ochsner 1928VC88
  N. olivaria Le Gillou 1841B01
  N. orbignyana Récluz 1841B01
  N. ornataEO64
  N. oualaniensisR43a
  N. panayensis Récluz 1843R43b
  N. pentastoma Deshayes 1866B01
  N. pfeifferiana Récluz 1843R43a
  ‘Neritina’ plutonis Basterot 1825B01
  N. polita Linnaeus 1758BK03, B01 (see below for synonymy)
  N. powisiana Récluz 1843R43a
  N. reticulata Karsten 1789 [incl. N. signata Lamarck 1822]B01
  N. royssianaR43a
  N. rumphii Récluz 1841R43b
  N. sanguinolenta Menke 1829 (see below for synonymy)BD86
  N. sayana Récluz 1843R43b
  N. sharonae (Squires & Saul 2001) [=Otostoma sharonae]BK03
  N. siquijorensis Récluz 1843R43b
  N. spengleriana Récluz 1843R43b
  N. squamulata Le Guillou 1841W93
  N. squiresi Bandel & Kiel 2003BK03
  N. subcompacta Trechmann 1927BK03
  N. textilisSP72
  N. tricarinataB01
  N. turtoni Récluz 1843R43a
  N. umzambiensis Wood 1906BK03
  N. variegata [incl. N. pulchra]R43a
  N. versicolor Gmelin 1791 [incl. N. versicolor f. nigrocincta Usticke 1959]BC01
  N. yoldiiF66

Nerita Linnaeus 1758VC88 [=Neritarius Duméril 1806KC60, Tenare Gray 1858KC60; incl. Dontostoma Herrmannsen 1847BK03, Linnerita Vermeij 1984B01]

Nerita polita Linnaeus 1758BK03, B01 [=*Dontostoma politaB01, N. (*Linnerita) politaB01; incl. N. bifasciata Gmelin 1791W93, N. litterata Gmelin 1791W93, N. vitiensis Hombron & Jacquinot 1854W93]

Nerita sanguinolenta Menke 1829 [incl. N. crassilabrum Smith 1885, N. forskalii Recluz 1841, N. kinzelbachi Nordsieck 1973]BD86

Nerita (*Cymostyla) undata Linnaeus 1758B01 [incl. N. chrysostoma Récluz 1841W93, N. flammulata Récluz 1841W93, N. leguillouana Récluz 1842W93, N. longii Récluz 1842W93, N. novaeguineae Lesson 1831W93, N. oleagina Reeve 1855W93, N. papuana Récluz 1841W93, N. quadricolor Gmelin 1791W93, N. savieana Reeve 1855W93, N. striata Burrow 1815W93, N. tongaensis Hombron & Jacquinot 1854W93, N. undulata Gmelin 1791W93]

Nerita (*Ilynerita) planospira Anton 1839VC88 [incl. N. angularis Hombron & Jacquinot 1854W93, Paludina angularisTC89, Vivipara angularisF66, Nerita atropurpurea Récluz 1841W93]

Nerita (Melanerita) atramentosa Reeve 1855B01 [incl. N. atrata Gmelin 1791W93, N. (*M.) nigra Martens 1889B01, N. saturata Hutton 1884W93]

Nerita (*Ritena) plicata Linnaeus 1758B01 [incl. N. eburnea Röding 1798W93, N. lactaria Linnaeus 1758W93, N. otaitensis Lesson 1831W93]

Nerita (*Theliostyla) albicilla Linnaeus 1758VC88 [=*Natere albicillaKC60; incl. N. imperfecta Röding 1798W93, N. ustulata Sowerby 1883W93, N. venusta Dunker 1844W93]

*Type species of generic name indicated


[B01] Bandel, K. 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen des Geologisch-Paläontologischen Institutes der Universität Hamburg 85: 65–164.

[BK03] Bandel, K., & S. Kiel. 2003. Relationships of Cretaceous Neritimorpha (Gastropoda, Mollusca), with the description of seven new species. Bulletin of the Czech Geological Survey 78 (1): 53–65.

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[EO64] Emerson, W. K., & W. E. Old Jr. 1964. Additional records from Cocos Island. Nautilus 77 (3): 90–92.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F66] Fraser, L. 1866. Communication of a list of mollusks collected by R. Swinhoe, Esq., in Formosa. Proceedings of the Zoological Society of London 1866: 146.

[G79] Guppy, R. J. L. 1879. First sketch of a marine invertebrate fauna of the Gulf of Paria and its neighbourhood. Part I.—Mollusca. Journal of Conchology 2: 151–172.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[O75] Owen, R. 1875. On fossil evidences of a sirenian mammal (Eotherium aegyptiacum, Owen) from the nummulitic Eocene of the Mokattam cliffs, near Cairo. Quarterly Journal of the Geological Society of London 31: 100–105.

[R42] Récluz, C. A. 1842. Descriptions of new species of nerites, collected by H. Cuming, Esq. in the Philippine Islands. Proceedings of the Zoological Society of London 10: 168–176.

[R43a] Récluz, C. A. 1843a. On new species of Nerita. Proceedings of the Zoological Society of London 11: 71–73.

[R43b] Récluz, C. A. 1843b. Descriptions of new species of Navicella, Neritina, Nerita and Natica, in the collection of Hugh Cuming, Esq. Proceedings of the Zoological Society of London 11: 197–214.

[SP72] Smythe, K. R., & W. W. A. Phillips. 1972. Some observations on the fauna of the Maldive Islands (Indian Ocean) part VIII. Marine shells. Journal of the Bombay Natural History Society 69 (2): 290–296.

[TC89] Tapparone Canefri, T. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XVIII.—Molluschi terrestri e d’acqua dolce. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 295–359.

[VC88] Vermeij, G. J., & T. M. Collins. 1988. Nerita fortidentata, a new gastropod from the Neogene of Panamá, with comments on the fossil record of Nerita in tropical America. Nautilus 102 (3): 102–105.

[W93] Wilson, B. 1993. Australian Marine Shells vol. 1. Prosobranch Gastropods. Part One. Odyssey Publishing: Kallaroo (Western Australia).

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

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