Nesticus cellulanus, copyright Gernot Kunz.

Belongs within: Araneoidea.

The Nesticidae, cave cobweb spiders, are a group of small spiders that primarily build their highly elastic cobwebs in dark locations. They are distinguished from the closely related Theridiidae by morphology of the male palp, in which the paracymbium is large and proximal, and the cymbium is rebordered (Jocqué & Dippenaar-Schoeman 2007).

The Synotaxidae are a group of small spiders found in Australasia and South America. They construct webs that vary from irregular sheets to an inverted bowl or lattice-like structure (Jocqué & Dippenaar-Schoeman 2007).

Characters (from Jocqué & Dippenaar-Schoeman 2007): Very small to small; eight eyes present in two rows; cheliceral fang furrow with two or three strong teeth and irregular group of numerous small denticles; labium rebordered; legs with three tarsal claws, tarsi IV with row of serrated bristles; ecribellate; entelegyne.

    |--Aituaria Esyunin & Efimik 1998JD-S07
    |--Canarionesticus Wunderlich 1992JD-S07
    |--Carpathonesticus Lehtinen & Saaristo 1980JD-S07
    |--Cyclocarcina Komatsu 1942JD-S07
    |--Gaucelmus Keyserling 1884JD-S07
    |--Typhlonesticus Kulczynski 1914JD-S07
    |--Baltonesticus flexuosus Wunderlich 1986S93
    |--Balticonesticus Wunderlich 1986P92
    |--Heteronesticus Wunderlich 1986P92
    |    `--H. magnoparacymbialis Wunderlich 1986S93
    |--Howaia Lehtinen & Saaristo 1980MG03
    |    `--*H. mogera (Yaginuma 1972) [=Nesticella mogera, Nesticus mogera]MG03
    |--Eidmannella Roewer 1935JD-S07
    |    `--E. rostrataJD-S07
    |--Nesticella Lehtinen & Saaristo 1980MG03
    |    |--*N. nepalensis (Hubert 1973)MG03
    |    |--N. robinsoni Lehtinen & Saaristo 1980MG03
    |    `--N. taurama Lehtinen & Saaristo 1980MG03
    |    |--E. inopinatus Wunderlich 1986S93
    |    |--E. longipes Petrunkevitch 1942S93
    |    `--E. rarus Wunderlich 1986S93
    `--Nesticus Thorell 1869 [Nesticeae]JD-S07
         |--N. akiensisT07
         |--N. akiyoshiensisT07
         |--N. cellulanus (Clerck 1757)GE02
         |--N. henderickxiJD-S07
         |--N. nasicusTH08
         |--N. noroensisT07
         `--N. silvestriiGE02

*Type species of generic name indicated


[GE02] Giribet, G., G. D. Edgecombe, W. C. Wheeler & C. Babbitt. 2002. Phylogeny and systematic position of Opiliones: a combined analysis of chelicerate relationships using morphological and molecular data. Cladistics 18: 5–70.

[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[MG03] Marusik, Yu. M., & E. F. Guseinov. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 1. New family and genus records. Arthropoda Selecta 12 (1): 29–46.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[TH08] Thomas, S. M., & M. Hedin. 2008. Multigenic phylogeographic divergence in the paleoendemic southern Appalachian opilionid Fumontana deprehendor Shear (Opiliones, Laniatores, Triaenonychidae). Molecular Phylogenetics and Evolution 46 (2): 645–658.

[T07] Tsurusaki, N. 2007. The Chugoku Mountains—a hotspot of geographical differentiation of species. Taxa 22: 3–14.

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