Novialoidea

Belongs within: Pterosauria.
Contains: Pterodactyloidea.

The Novialoidea are a clade of pterosaurs defined by Kellner (2003) as the most exclusive clade containing Campylognathoides and Quetzalcoatlus. Members of this clade have the third phalanx of the wing finger shorter than the first and second phalanges (Kellner 2003). The exclusive clade Breviquartossa was defined to include Rhamphorhynchus and Quetzalcoatlus by Unwin (2003).

<==Novialoidea (see below for synonymy)K03
    |--Campylognathoides Strand 1928RK13, U03
    |    |--C. indicus Jain 1974W91
    |    |--C. liasicus (Quenstedt 1858)K03 [=Pterodactylus liasicusW13]
    |    `--C. zitteli [=Campylognathus zitteli]W13
    `--BreviquartossaU03
         |--PterodactyloideaRK13
         `--Rhamphorhynchus von Meyer 1847RK13, A08
              |--R. jessoni Lydekker 1890W91
              |--R. longicaudus (Münster 1839)RK13, K03 [=Pterodactylus longicaudusW91]
              |--R. longiceps Woodward 1902B93 [incl. R. kokeniW91]
              |--R. muensteri (Goldfuss 1831)W13 (see below for synonymy)
              |--R. phyllurus Marsh 1882W91
              `--R. tendagurensis Reck 1931W91
Novialoidea incertae sedis:
  Carniadactylus Dalla Vecchia 2009W13
    `--*C. rosenfeldi (Dalla Vecchia 1995)W13, DV03 [=Eudimorphodon rosenfeldiW13]
  Fenghuangopterus liiW13
  Jianchangopterus zhaoianusW13
  Pterorhynchus Czerkas & Ji 2002RK13, A08
    `--P. wellnhoferiW13
  Sericipterus wucaiwanensisW13
  Qinglongopterus guoiW13
  Bellubrunnus rothgaengeriW13
  Nesodactylus Colbert 1969U03
    `--N. hesperius Colbert 1969B93
  Rhamphocephalus Seeley 1880 [incl. Dolichorhamphus Seeley 1885, Pteromonodactylus Teryaev 1967]A08
    |--R. bucklandi (von Meyer 1832) [=Pterodactylus bucklandi]W91
    |--R. depressirostrisW13
    `--R. prestwichiW91
  Cacibupteryx Gasparini, Fernández & de la Fuente 2004RK13, A08
    `--*C. caribensis Gasparini, Fernández & de la Fuente 2004GFF04
  Dorygnathus Wagner 1860U03 [incl. Parapsicephalus Arthaber 1919W13]
    |--D. banthensis (Theodori 1830)K03 (see below for synonymy)
    `--D. purdoni Newton 1888 [=Scaphognathus purdoni, Parapsicephalus purdoni]W13
  Angustinaripterus He, Yan & Su 1983U03
    `--A. longicephalusRK13
  Harpactognathus Carpenter, Unwin et al. 2003CU03
    `--*H. gentryii Carpenter, Unwin et al. 2003CU03
Inorganic: Rhamphorhynchus gemmingi minilorientalus Okamura 1987O87

Dorygnathus banthensis (Theodori 1830)^K03 [=Ornithocephalus banthensis^W91, Pterodactylus banthensis^W91; incl. Dorygnathus mistelgauensis^W13, Parapsicephalus mistelgauensis^D07]

Novialoidea [Angustinaripterinae, Campylognathoididae, Monofenestrata, Parapsicephalinae, Pteromonodactylidae, Rhamphorhynchae, Rhamphorhynchinae]^K03

Rhamphorhynchus muensteri (Goldfuss 1831)^W13 [=Ornithocephalus muensteri^W13, Pterodactylus muensteri^W91; incl. R. gemmingi Meyer 1846^{U03, B93}, R. intermedius Koh 1937^{U03, B93}]

*Type species of generic name indicated

References

[A08] Averianov, A. O. 2008. Superorder Pterosauria. In: Ivakhnenko, M. F., & E. N. Kurochkin (eds) Fossil Vertebrates of Russia and Adjacent Territories. Fossil Reptiles and Birds. Part 1 pp. 319–342. GEOS: Moscow.

[B93] Benton, M. J. 1993. Reptilia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 681–715. Chapman & Hall: London.

[CU03] Carpenter, K., D. Unwin, K. Cloward, C. Miles & C. Miles. 2003. A new scaphognathine pterosaur from the Upper Jurassic Morrison Formation of Wyoming, USA. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 45–54. The Geological Society: London.

[DV03] Dalla Vecchia, F. M. 2003. New morphological observations on Triassic pterosaurs. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 23–44. The Geological Society: London.

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[GFF04] Gasparini, Z., M. Fernández & M. de la Fuente. 2004. A new pterosaur from the Jurassic of Cuba. Palaeontology 47: 919–927.

[K03] Kellner, A. W. A. 2003. Pterosaur phylogeny and comments on the evolutionary history of the group. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 105–137. The Geological Society: London.

[O87] Okamura, C. 1987. New facts: Homo and all Vertebrata were born simultaneously in the former Paleozoic in Japan. Original Report of the Okamura Fossil Laboratory 15: 347–573.

[RK13] Rodrigues, T., & A. W. A. Kellner. 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308: 1–112.

[U03] Unwin, D. M. 2003. On the phylogeny and evolutionary history of pterosaurs. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 139–190. The Geological Society: London.

[W91] Wellnhofer, P. 1991. The Illustrated Encyclopedia of Pterosaurs.  Salamander Books: London (reprinted in: Norman, D., & P. Wellnhofer. 2000. The Illustrated Encyclopedia of Dinosaurs. Salamander Books).

[W13] Witton, M. P. 2013. Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press: Princeton (New Jersey).