Nyssonini

Metanysson coahuila, copyright CBG Photography Group.

Belongs within: Bembicidae.
Contains: Epinysson, Zanysson, Nysson, Synnevrus, Brachystegus.

The Nyssonini are a group of medium-sized to small wasps, usually with a heavily armoured integument, that are kleptoparasites of other Bembicidae or Crabronidae (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Eyes converging below, often strongly, inner margins slightly to moderately emarginate above; ocelli well developed. Labrum small to rather large, in latter case broadly subrectanpular; palpal formula 6-4; pronotum closely appressed to scutum. Scutum with admedian lines fused into a single line at or very near anterior edge; notauli absent; with oblique scutal carina; scutellum without posterior overlapping edge. Omaulus usually present, sometimes indistinct; acetabular carina present, sometimes faint; subomaulus present; episternal and scrobal sulci absent or evanescent; sternaulus absent; verticaulus sometimes present. Metapleuron not always separated from propodeum by a distinct sulcus or line beneath upper metapleural pit; propodeum usually spined dorsolaterally. Midcoxae widely separated, precoxal sulcus complete or sometimes obscure; midtibia usually with two apical spurs (exceptionally one); foretarsal rake weak or absent; arolia small or absent; plantulae present but usually very small. Forewing media usually diverging at or before cu-a; stigma usually rather small; two or three submarginal cells, true second cell petiolate (when present) and receiving at least one recurrent vein. Hindwing media diverging before to after cu-a; jugal lobe small or absent. Gaster with tergum I nearly as broad as II and with lateral line or carina; sternum I with a double ridge converging basally; male with seven visible terga; male sternum VIII simple; male genitalia with volsella undifferentiated into cuspis and digitus; female pygidial plate usually well developed but sometimes indistinct, and with entire pygidium serrate or multidentate.

<==NyssoniniBM76
| i. s.: EpinyssonBM76
| ZanyssonBM76
| Neonysson Bohart 1968BM76
| |--*N. porteri (Ruiz 1936) [=Nysson porteri]BM76
| `--N. herbsti Bohart 1968BM76
| Hovanysson Arnold 1945BM76
| |--*H. camelus (Arnold 1945) [=Brachystegus camelus]BM76
| `--H. albibarbis (Arnold 1945) [=Nysson albibarbis]BM76
|--+--Nursea Cameron 1902BM76
| | `--*N. carinata Cameron 1902BM76
| `--Nippononysson Yasumatsu & Maidl 1936BM76
| |--*N. rufopictus Yasumatsu & Maidl 1936BM76
| |--N. adiaphilus Krombein 1943BM76
| `--N. inexspectatus Beaumont 1967BM76
`--+--+--NyssonBM76
| |--SynnevrusBM76
| `--Hyponysson Cresson 1882BM76
| |--*H. bicolor Cresson 1882BM76
| `--H. raui (Rohwer 1917) [=Nysson raui]BM76
`--+--+--+--BrachystegusBM76
| | `--Heterogyna [Heterogynaidae]SN18
| | |--H. madecassaB11
| | |--H. nocticolaSN18
| | `--H. proteaB11
| `--Acanthostethus Smith 1869 [incl. Spalagia Shuckard 1840 (n. n.)]BM76
| |--A. mysticus (Gerstäcker 1867) [=Nysson mysticus; incl. *A. basalis Smith 1869]C85
| |--A. brisbanenssi (Turner 1915) [=Nysson (Acanthostethus) brisbanensis]C85
| |--A. confertus (Turner 1915) [=Nysson (Acanthostethus) confertus]C85
| |--A. gilberti (Turner 1915) [=Nysson (Acanthostethus) gilberti]C85
| |--A. hentyi (Rayment 1953) [=Nysson hentyi]C85
| |--A. minimus (Turner 1915) [=Nysson (Acanthostethus) minimus]C85
| |--A. moerens (Turner 1915) [=Nysson (Acanthostethus) moerens]C85
| |--A. nudiventris (Turner 1915) [=Nysson (Acanthostethus) nudiventris]C85
| |--A. obliteratus (Turner 1910) [=Nysson (Acanthostethus) obliteratus]C85
| |--A. portlandensis (Rayment 1953) [=Nysson portlandensis]C85
| |--A. punctatissimus (Turner 1908) [=Nysson (Acanthostethus) punctatissimus]C85
| |--A. saussurei (Handlirsch 1887) [=Nysson saussurei]C85
| |--A. spiniger (Turner 1908) [=Nysson (Acanthostethus) spiniger]C85
| |--A. tasmanicus (Turner 1915) [=Nysson (Acanthostethus) tasmanicus]C85
| `--A. triangularis (Turner 1940) [=Nysson (Acanthostethus) triangularis]C85
`--+--+--Perisson Pate 1938BM76
| | `--*P. basirufus (Brèthes 1913) [=Nysson basirufus]BM76
| |--Cresson Pate 1938 [=Cressonius Bradley 1956]BM76
| | `--*C. parvispinosus (Reed 1894) [=Nysson parvispinosus]BM76
| |--Antomartinezius Fritz 1955BM76
| | |--*A. patei (Fritz 1955) [=Cresson patei]BM76
| | |--A. boharti Fritz 1973BM76
| | `--A. fritzi Bohart 1968BM76
| `--Foxia Ashmead 1898BM76
| |--*F. pacifica Ashmead 1898BM76
| |--F. cuna Pate 1938BM76
| |--F. deserticola Fritz 1959BM76
| |--F. divergens (Ducke 1903) [=Nysson divergens]BM76
| |--F. garciai Fritz 1972BM76
| |--F. martinezi Fritz 1972BM76
| |--F. navajo Pate 1938BM76
| |--F. pirita Fritz 1972BM76
| |--F. secunda (Rohwer 1921) [=Nysson secundus]BM76
| `--F. tercera Fritz 1972BM76
`--+--Losada Pate 1940BM76
| |--*L. paria Pate 1940BM76
| |--L. mutilloides (Ducke 1903) [=Nysson mutilloides]BM76
| `--L. penai Fritz 1973BM76
|--Idionysson Pate 1940BM76
| |--*I. borero Pate 1940BM76
| |--I. chrysozonus (Gerstaecker 1867) [=Nysson chrysozonus]BM76
| `--I. cordialis Fritz 1970BM76
`--Metanysson Ashmead 1899 [incl. Huachuca Pate 1938]BM76
|--*M. solani (Cockerell 1895) [=Nysson solani]BM76
|--M. alfkeni (Ducke 1904) [=Nysson alfkeni]BM76
|--M. arivaipa Pate 1938 [=*Huachuca arivaipa]BM76
|--M. carcavalloi Fritz 1959BM76
|--M. catamarcensis (Schrottky 1910) [=Paranysson catamarcensis]BM76
|--M. cicheroi Fritz 1973BM76
|--M. coahuila Pate 1938BM76
|--M. diezguitas Fritz 1958BM76
|--M. foersteri Fritz 1958BM76
|--M. fraternus Fritz 1970BM76
|--M. horacioi Fritz 1973BM76
|--M. layano Pate 1938BM76
|--M. lipan Pate 1938BM76
|--M. toba Fritz 1973BM76
|--M. tropicalis Fritz 1970BM76
`--M. yavapai Pate 1938BM76

*Type species of generic name indicated

References

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[B11] Brothers, D. J. 2011. A new Late Cretaceous family of Hymenoptera, and phylogeny of the Plumariidae and Chrysidoidea (Aculeata). ZooKeys 130: 515–542.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

[SN18] Sann, M., O. Niehuis, R. S. Peters, C. Mayer, A. Kozlov, L. Podsiadlowski, S. Bank, K. Meusemann, B. Misof, C. Bleidorn & M. Ohl. 2018. Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. BMC Evolutionary Biology 18: 71.

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