Ocypodoidea

Horned ghost crab Ocypode ceratophthalma, copyright Adrien Chateignier.

Belongs within: Brachyura.
Contains: Uca.

The Ocypodoidea are a group of crabs primarily found in sandy intertidal habitats, many of which are characterised by elongate eye-stalks.

<==OcypodoideaMD01
    |--Camptandriidae [Camptandriinae]MD01
    |--MictyrisT03 [MictyridaeMD01]
    |    |--M. brevidactylus Stimpson 1858T03
    |    `--M. longicarpus Latreille 1806J90 [=Leucosia (Mictyris) longicarpaG20]
    |--PalicidaeMD01
    |    |--Micropalicus vietnamensis (Zarenkov 1968)MG-H11
    |    |--Neopalicus jukesi (White 1847)MG-H11
    |    |--Paliculus kyusyuensis (Yokoya 1933)MG-H11
    |    |--ParapalicusMG-H11
    |    `--Pseudopalicus macromeles Castro 2000MG-H11
    `--OcypodidaeB68
         |  i. s.: Hemiplax Heller 1865B64
         |           |--H. hirtipes (Jacquinot in Jacquinot & Lucas 1853) (see below for synonymy)B64
         |           `--H. latifonsF71
         |         Loerentheya Beurlen 1929BWW93
         |           `--L. carinata Beurlen 1929BWW93
         |--DotillinaeMD01
         |--ScopimerinaeB68
         |    |--IlyoplaxB68
         |    |--Tmethypocoelis ceratophoraB68
         |    `--Scopimera inflataCJF98
         |--Macropthalminae [Macrophthalmidae]B68
         |    |--AustraloplaxB68
         |    `--MacrophthalmusB68
         |         |--M. (Chaenostoma) boscii Audouin 1826TSH09
         |         |--M. depressusF71
         |         |--M. graeffei Milne-Edwards 1873KK03
         |         |--M. hirtipesHS01
         |         |--M. serratusF71
         |         |--M. setosusF71
         |         `--M. verreauxi Milne Edwards 1848J90
         `--OcypodinaeB68
              |--UcaB68
              |--Heloecius [Heloeciinae]B68
              |    `--H. cordiformisB64
              `--Ocypode Fabricius 1798B55
                   |--O. ceratophthalma (Pallas 1772) [incl. O. urvillei Guerin 1829]ST76
                   |--O. convexa Quoy & Gaimard 1824J90
                   |--O. cordimana Latreille 1818ST76
                   |--O. cursor (Linnaeus 1758)KK03
                   |--O. fabricii Milne Edwards 1837J90
                   |--O. gaudichaudii Milne Edwards & Lucas 1843PTB03 [=Ocypoda gaudichaudiiB55]
                   |--O. kuhlii de Haan 1835ST76
                   |--O. macrocera Milne-Edwards 1837 [incl. Ocypoda portonovoensis Premkumar 1964]ST76
                   |--O. occidentalis Stimpson 1860 [=Ocypoda occidentalis]B55
                   |--O. pallidula Jacquinot in Hombron & Jacquinot 1842-1847 [incl. Ocypoda laevis Dana 1852]ST76
                   |--O. quadrata (Fabricius 1787)PTB03 [incl. O. albicans Bosc 1802PTB03, O. arenariaST76]
                   |--O. ruricola (Linnaeus 1758)G20, L58 [=Cancer ruricolaL02, Grapsus (Ocypode) ruricolaG20]
                   |--O. ryderi Kingsley 1881ST76
                   `--O. vocans (Linnaeus 1758)L02, L58 [=Cancer vocansL02]

Hemiplax hirtipes (Jacquinot in Jacquinot & Lucas 1853) [=Cleistostoma hirtipes; incl. Metaplax hirtipes Heller 1862 non Cleistostoma hirtipes Jacquinot in Jacquinot & Lucas 1853]B64

*Type species of generic name indicated

References

[B68] Barnes, R. S. K. 1968. On the evolution of elongate ocular peduncles by the Brachyura. Systematic Zoology 17 (2): 182–187.

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[B55] Bott, R. 1955. Dekapoden (Crustacea) aus El Salvador. 2. Litorale Dekapoden, außer Uca. Senckenbergiana Biologica 36: 45–70.

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[CJF98] Casanova, J.-P., L. D. Jong & E. Faure. 1998. Interrelationships of the two families constituting the Lophogastrida (Crustacea: Mysidacea) inferred from morphological and molecular data. Marine Biology 132: 59–65.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.

[J90] Jones, D. S. 1990. Annotated checklist of marine decapod Crustacea from Shark Bay, Western Australia. In: Berry, P. F., S. D. Bradshaw & B. R. Wilson (eds) Research in Shark Bay: Report of the France-Australe Bicentenary Expedition Committee pp. 169–208. Western Australian Museum.

[KK03] Kocataş, A. & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[PTB03] Portell, R. W., R. L. Turner & J. L. Beerensson. 2003. Occurrence of the Atlantic ghost crab Ocypode quadrata from the Upper Pleistocene to Holocene Anastasia Formation of Florida. Journal of Crustacean Biology 23 (3): 712–722.

[ST76] Sakai, K., & M. Türkay. 1976. Bemerkungen zu einigen Ocypode-Arten (Crustacea: Decapoda). Senckenbergiana Biologica 57 (1–3): 81–96.

[T03] Takeda, S. 2003. Mass wandering in the reproductive season by the fiddler crab Uca perplexa (Decapoda: Ocypodidae). Journal of Crustacean Biology 23 (3): 723–728.

[TSH09] Titelius, M. A., A. Sampey & C. G. Hass. 2009. Crustaceans of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, north Western Australia. Records of the Western Australian Museum Supplement 77: 145–176.

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