Omaliinae

Omalium rivulare, copyright Udo Schmidt.

Belongs within: Staphylinidae.
Contains: Coryphiina.

The Omaliinae are a group of staphylinid beetles typically possessing a pair of ocelli near the hind margin of the head (Newton et al. 2001). The subfamily as historically recognised is resolved by molecular data as paraphyletic to taxa that have been assigned to separate subfamilies Glypholomatinae, Microsilpha and Empelus (Lü et al. 2020).

<==Omaliinae [Anthophagini, Brathinidae, Glypholomatinae, Omalides, Omaliini]
|--+--Olophrum Erichson 1839LC20, NT01 [incl. Lathrium LeConte 1850NT01]
| | |--O. fuscumN90
| | |--O. obtectum Erichson 1840NT01
| | `--O. piceumLC20
| `--Phyllodrepa Thomson 1859LC20, NT01
| |--P. devilleiLC20
| `--P. floralis (Paykull 1789)NT01
`--+--+--Anthobiomimus unicolorLC20
| `--+--+--Eusphalerum Kraatz 1858LC20, NT01 [incl. Onibathum Tottenham 1939NT01; Eusphalerini]
| | | `--E. torquatum (Marsham 1802)NT01
| | `--Amphichroum Kraatz 1858LC20, NT01 [incl. Stachygraphis Horn 1883NT01]
| | |--A. adelaidae Blackb. 1892M96
| | |--A. floribundum LeConte 1863NT01
| | |--A. maculatum Horn 1882NT01
| | `--A. maculicolle (Mannerheim 1843)NT01
| `--+--Acrolocha Thomson 1858LC20, NT01 [incl. Elonium Leach 1819 (nom. rej.)NT01]
| | |--A. diffusa (Fauvel 1878)NT01
| | |--A. helferi Steel 1957NT01
| | |--A. leechi Hatch 1957NT01
| | |--A. minutaN90
| | |--A. newtoni Thayer 2003GN09
| | `--A. sulculaLC20
| `--+--Omaliopsis Jeannel 1940LC20, NT01
| | |--O. ectopiaLC20
| | `--O. russataNT01
| `--Omalium Gravenhorst 1802LC20, NT01 [=Homalium Ljungh 1804NT01]
| |--O. adelaidae Blackb. 1887M96
| |--O. caesumLC20
| |--O. florale [=Oxytelus (Omalium) floralis]G20
| |--O. fractum Fauvel 1878NT01
| |--‘Homalium’ lachrymale Fauv. in Fleutiaux & Sallé 1890FS90
| |--O. punctatumR26
| `--O. rivulareN90
`--+--Metacorneolabium convexumLC20
`--+--+--Proglypholoma aenigmaLC20
| `--Microsilpha [Microsilphinae, Microsilphini]LC20
| `--M. ocelligera (Champion 1918)B14
`--+--GlypholomaLC20
| |--G. peckiLC20
| `--G. rotundulum Thayer & Newton 1979B14
`--+--+--Pelecomalium Casey 1886LC20, NT01 [incl. Heterops Mannerheim 1843 non Blanchard 1842NT01]
| | `--P. flavescensNT01
| `--Brathinus LeConte 1852LC20, NT01
| |--B. californicus Hubbard 1894NT01
| |--B. nitidus LeConte 1852NT01
| `--B. varicornis LeConte 1852NT01
`--+--+--Anthobium Leach 1819LC20, NT01 [incl. Deliphrum Erichson 1839NT01, Lathrimaeum Erichson 1839NT01]
| | |--A. minutumC01
| | `--A. unicolorN90
| `--Arpedium Erichson 1839LC20, NT01
| |--A. angulare Fauvel 1878NT01
| |--A. brachypterumN90
| |--A. cribratum Fauvel 1878NT01
| |--A. quadrum (Gravenhorst 1806)NT01
| `--A. schwarzi Fauvel 1878NT01
`--+--Empelus LeConte 1861LC20, NT01 [Empelidae, Empelinae]
| `--E. brunnipennis (Mannerheim 1852) [=Litochrus brunnipennis]B14
`--+--Lesteva Latreille 1796LC20, NT01 (see below for synonymy)
| |--*L. abbreviata [=Carabus abbreviatus]L02
| |--L. alpinaR26
| |--L. cribratula (Casey 1893) [=Tevales cribratulus]NT01
| |--L. pallipes LeConte 1863 [=Pseudolesteva pallipes]NT01
| `--L. punctulataR26
`--CoryphiiniNT01
|--CoryphiinaLC20
`--BoreaphilinaNT01
|--Boreaphilus Sahlberg 1832NT01
| `--B. henningianus Sahlberg 1832NT01
`--Gnathoryphium Campbell 1978NT01
`--G. mandibulare Campbell 1978NT01

Omaliinae incertae sedis:
Sinanthobium daohugouensisLC20
HadrognathiniNT01
CorneolabiiniNT01
AphaenostemminiNT01
|--Aphaenostemmus Peyerimhoff 1914NT01
`--Giulianium Moore 1976NT01
|--G. alaskanum Ahn & Ashe 1999NT01
|--G. campbelli Moore 1976NT01
`--G. newtoni Ahn & Ashe 1999NT01
Omalonomus Campbell & Peck 1990NT01
`--O. relictus Campbell & Peck 1990NT01
Paraphloeostiba Steel 1960NT01
`--P. gayndahensis (MacLeay 1871)NT01
Phloeostiba Thomson 1858 [incl. Distemmus LeConte 1861]NT01
`--P. lapponicus (Zetterstedt 1838)NT01
Acrulia Thomson 1858NT01
|--A. inflataN90
`--A. tumidula (Mäklin 1853) [incl. Acrolocha crenulata Hatch 1957]NT01
Carcinocephalus Bernhauer 1903NT01
|--C. exsculptus (Mäklin 1852) [=Omalium exsculptum; incl. Acrolocha rugosa Hatch 1957]NT01
`--C. flavidus (Hamilton 1895) [=Omalium flavidum]NT01
Dropephylla Mulsant & Rey 1880NT01
|--D. cacti (Schwarz 1899)NT01
`--D. longula (Mäklin 1852) [=Hapalaraea longula]NT01
Micralymma Westwood 1838NT01
|--M. brevilingue Schiødte 1845NT01
`--M. marinum (Ström 1783)NT01
Xylodromus Heer 1839NT01
|--X. capito Casey 1893NT01
|--X. concinnus (Marsham 1802)NT01
`--X. depressus (Gravenhorst 1802)NT01
Hapalaraea Thomson 1858NT01
|--H. hamata (Fauvel 1878) [incl. Acrolocha barri Hatch 1957]NT01
|--H. megarthroides (Fauvel 1878)NT01
`--H. pygmaeaN90
Phloeonomus Heer 1839NT01
|--P. laesicollis Mäklin 1852NT01
|--P. pusillus (Gravenhorst 1806)NT01
`--P. suffusus Casey 1893NT01
Pycnoglypta Thomson 1858NT01
|--P. aptera Campbell 1983NT01
|--P. campbelli Gusarov 1995NT01
|--P. heydeni Eppelsheim 1886NT01
`--P. lurida (Gyllenhal 1813)NT01
Mannerheimia Mäklin 1880NT01
Phlaeopterus Motschulsky 1853 [incl. Tilea Fauvel 1878]NT01
Unamis Casey 1893NT01
Anthophagus Gravenhorst 1802NT01
`--A. caraboides [=Oxytelus (Anthophagus) caraboides]G20
Phyllodrepoidea Ganglbauer 1895NT01
`--P. crenata (Gravenhorst 1802)NT01
Anthobioides Campbell 1987NT01
`--A. pubescens Campbell 1987NT01
Geodromicus Redtenbacher 1856 [incl. Psephidonus Gistel 1856]NT01
`--G. brunneusNT01
Microedus LeConte 1874NT01
`--M. austinianus LeConte 1874NT01
Omalorphanus Campbell & Chandler 1987NT01
`--O. aenigma Campbell & Chandler 1987NT01
Orobanus LeConte 1878NT01
`--O. densusNT01
Tanyrhinus Mannerheim 1852NT01
`--T. singularis Mannerheim 1852B14
Vellica Casey 1885NT01
`--V. longipennis Casey 1885NT01
Xenicopoda Moore & Legner 1971NT01
`--X. helenae Moore & Legner 1971NT01
Artochia Casey 1893NT01
|--A. californica Bernhauer 1912NT01
`--A. productifrons Casey 1893NT01
Porrhodites Kraatz 1858NT01
|--P. fenestralis (Zetterstedt 1828)NT01
`--P. inflatus (Hatch 1957)NT01
Trigonodemus LeConte 1863NT01
|--T. fasciatus Leech 1939NT01
`--T. striatus LeConte 1863NT01
Deinopteroloma Jansson 1946NT01
|--D. pictum (Fauvel 1878) [=Lathrimaeum pictum]NT01
|--D. spectabile Smetana 1985B14
`--D. subcostatum (Mäklin 1852) [=Lathrimaeum subcostatum]NT01
Eucnecosum Reitter 1909 [incl. Revelstokea Hatch 1957]NT01
|--E. brachypterum (Gravenhorst 1802)NT01
|--E. brunnescens (Sahlberg 1871)NT01
`--E. tenue (LeConte 1863)NT01
Orochares Kraatz 1858 [incl. Paradeliphrum Hatch 1957]NT01
|--O. angustatus (Erichson 1839)NT01
|--O. suteri Campbell 1984NT01
`--O. tumidus (Hatch 1957)NT01
Acidota Stephens 1829NT01
|--A. crenata (Fabricius 1792)NT01
|--A. cruentata Mannerheim 1830NT01
|--A. quadrata (Zetterstedt 1838)NT01
`--A. subcarinata Erichson 1839NT01
Juroglypholoma antiquumMW15

Nomen nudum: Homalium arpedinum Fauvel in Keen 1895NT01

Lesteva Latreille 1796LC20, NT01 [incl. Lesta Blackwelder 1952NT01, Paralesteva Casey 1905NT01, Pseudolesteva Casey 1893NT01, Tevales Casey 1893NT01]

*Type species of generic name indicated

References

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[GN09] Grebennikov, V. V., & A. F. Newton. 2009. Good-bye Scydmaenidae, or why the ant-like stone beetles should become megadiverse Staphylinidae sensu latissimo (Coleoptera). European Journal of Entomology 106: 275–301.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LC20] Lü, L., C.-Y. Cai, X. Zhang, A. F. Newton, M. K. Thayer & H.-Z. Zhou. 2020. Linking evolutionary mode to palaeoclimate change reveals rapid radiations of staphylinoid beetles in low-energy conditions. Current Zoology 66 (4): 435–444.

[M96] Masters, G. 1896. Catalogue of the described Coleoptera of Australia. Supplement, part II. Dytiscidae, Gyrinidae, Hydrophyllidae, Staphylinidae, Pselaphidae, Paussidae, Silphidae, Scaphididae, Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydiidae, Cucujidae, Cryptophagidae, Lathridiidae, Mycetophagidae, Dermestidae, Byrrhidae, Parnidae, Heteroceridae. Proceedings of the Linnean Society of New South Wales 21 (Suppl.): 695–754.

[N90] Newton, A. F., Jr. 1990. Insecta: Coleoptera Staphylinidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1137–1174. John Wiley & Sones: New York.

[NT01] Newton, A. F., M. K. Thayer, J. S. Ashe & D. S. Chandler. 2001. Staphylinidae Latreille, 1802. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 272–418. CRC Press: Boca Raton.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

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