Ophiodermatidae

Ophiopeza spinosa, copyright Moorea Biocode.

Belongs within: Ophiurida.
Contains: Ophiocomidae.

The Ophiodermatidae are a group of brittle stars with the relatively stout arms inserted laterally and firmly fused with the disc (Spencer & Wright 1966).

Characters (from Spencer & Wright 1966): Arms inserted laterally and firmly fused with disc, moderately long, stout, widest at base. Granules cover disc scales of both surfaces and commonly jaws also. Unpaired infradental papillae at apex of each jaw.

<==Ophiodermatidae [Ophiarachnidae]
    |--+--OphiocomidaeTS16
    |  `--Ophiarachna Müller & Troschel 1842TS16, SW66
    |       |--*O. incrassata (Lamarck 1816) [=Ophiura incrassata]SW66
    |       |--O. affinis Lütken 1869SM15
    |       `--O. delicata (Clark 1932)SM15
    `--Ophioderma Müller & Troschel 1840TS16, SW66
         |--*O. longicauda (Bruzelius 1805)TS16 [=Asterias longicaudaSW66]
         |--O. asteriformisB89
         |--O. brevispinumF67
         |--O. egertoniB89
         |--O. gareyiB89
         `--O. hauchecornei [=Ophiarachna hauchecornei]B89
Ophiodermatidae incertae sedis:
  Cryptopelta Clark 1909 [incl. Ophiodyscrita Clark 1938]SW66
    |--*C. aster (Lyman 1879) [=Ophiopeza aster]SW66
    |--‘Ophiodyscrita’ acosmeta Clark 1938SM15
    |--C. callista Clark 1938SM15
    `--C. granulifera Clark 1909SM15
  Ophiochasma Grube 1868 [incl. Ophiopinax Bell 1884]SW66
    `--O. stellata (Ljungman 1867) [=Ophiarachna stellata; incl. *Ophiochasma adspersa Grube 1868]SW66
  Ophiopsammus Lütken 1869MG-H11, SW66
    `--O. assimilis (Bell 1888)MG-H11
  Bathypectinura Clark 1909SW66
    `--*B. lacertosa (Lyman 1883) [=Pectinura lacertosa]SW66
  Diopederma Clark 1913SW66
    `--*D. daniana (Verrill 1867) [=Ophiura daniana]SW66
  Distichophis Ely 1942SW66
    `--*D. clarki Ely 1942SW66
  Ophiochaeta Lütken 1869SW66
    `--*O. hirsuta Lütken 1869SW66
  Ophioclastus Murakami 1943SW66
    `--*O. hataii Murakami 1943SW66
  Ophiocormus Clark 1915 [incl. Ophiostegastus Murakami 1944]SW66
    `--*O. notabilis Clark 1915SW66
  Ophiocryptus Clark 1915SW66
    `--*O. maculosus Clark 1915SW66
  Ophioncus Ives 1889SW66
    `--*O. granulosus Ives 1889SW66
  Ophiopaepale Ljungman 1871SW66
    `--*O. goesiana Ljungman 1871SW66
  Ophiopetra Hess 1962SW66
    `--*O. lithographica Hess 1962SW66
  Ophiopezella Ljungman 1871SW66
    `--*O. spinosa (Ljungman 1867) [=Ophiarachna spinosa]SW66
  Ophiurochaeta Matsumoto 1915SW66
    `--*O. mixta (Lyman 1878) [=Ophiochaeta mixta]SW66
  Ophiuroconis Matsumoto 1915SW66
    `--*O. monolepis Matsumoto 1915SW66
  Ophiurodon Matsumoto 1915SW66
    `--*O. grandisquama (Koehler 1904) [=Ophioconis grandisquama]SW66
  Schizoderma Nielsen 1932 nec Ehrenb. 1818 (ICBN) nec Kunze 1825 (ICBN)SW66
    `--*S. diplax Nielsen 1932SW66
  Toporkovia Djakonov 1954SW66
    `--*T. fragilis Djakonov 1954SW66
  Pectinura Forbes 1843SW66
    |--*P. vestita Forbes 1843SW66
    `--P. cylindricaF67
  Ophioconis Lütken 1869SW66
    |--*O. forbesi (Heller 1862) [=Pectinura forbesi]SW66
    |--O. cincta Brock 1888SM15
    `--O. opacum (Clark 1928)MG-H11
  Ophiarachnella Ljungman 1872SW66
    |--*O. gorgonia (Müller & Troschel 1842) [=Ophiarachna gorgonia]SW66
    |--O. infernalis (Müller & Troschel 1842)SM15
    |--O. septemspinosa (Müller & Troschel 1842)BM09
    |--O. snelliusi (Clark 1964)SM15
    `--O. sphenisci (Bell 1894)SM15
  Ophiopeza Peters 1851SW66
    |--*O. fallax Peters 1851SW66
    |--O. cylindrica (Hutton 1872)MG-H11
    `--O. spinosa (Ljungman 1867)MG-H11

*Type species of generic name indicated

References

[B89] Boehm, G. 1889. Ein Beitrag zur Kenntniss fossiler Ophiuren. Berichte der Naturforschenden Gesellschaft zu Freiburg I. B. 4: 232–287, pls 4–5.

[BM09] Bryce, C., & L. Marsh. 2009. Echinodermata (Asteroidea, Echinoidea and Holothuroidea) of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 209–220.

[F67] Fell, H. B. 1967. Echinoderm ontogeny. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt S. Echinodermata 1. General characters. Homalozoa—Crinozoa (except Crinoidea) vol. 1 pp. S60–S85. The Geological Society of America, Inc., and The University of Kansas: Lawrence (Kansas).

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[SM15] Sampey, A., & L. M. Marsh. 2015. Kimberley marine biota. Historical data: echinoderms. Records of the Western Australian Museum Supplement 84: 207–246.

[SW66] Spencer, W. K., & C. W. Wright. 1966. Asterozoans. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt U. Echinodermata 3 vol. 1 pp. U4–U107. The Geological Society of America, Inc., and The University of Kansas Press.

[TS16] Thuy, B., & S. Stöhr. 2016. A new morphological phylogeny of the Ophiuroidea (Echinodermata) accords with molecular evidence and renders microfossils accessible for cladistics. PLoS One 11 (5): e0156140.

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