Porismus strigatus, copyright Donald Hobern.

Belongs within: Euneuroptera.

The Osmylidae are a diverse group of moderate-sized to large lacewings that typically have camouflaging patterns on their wings. Larvae are long and slender with elongate, lance-like jaws and are most commonly found in moist riparian habitats (Engel et al. 2018).

Characters (from New 1991): Generally medium to large (fore wing length 12-30 mm or more), with wings often patterned with grey or brown and rather broad; antennae usually less than half as long as fore wing; three ocelli present, at least represented by raised tubercles on vertex; female genitalia with enlarged gonocoxite 9, and 2 spermathecae. Larvae long and fusiform, with long tapering straight jaws.

    |    |--GumillaA69 [GumillinaeEWB18]
    |    |--SpilosmylinaeRY03
    |    |    |--Spilosmylus flavicornisRD77
    |    |    `--Conchylosmylus triseriatusR70b
    |    `--ProtosmylinaeEWB18
    |         |--Paryphosmylus ornatusA69
    |         `--ProtosmylusP92
         |  i. s.: EidoporismusN91 [EidoporisminaeEWB18]
         |           `--E. pulchellusN91
         |    |--PlethosmylusA69
         |    `--Osmylus Latreille 1802WHW10, L02
         |         |--*O. maculatus [=Hemerobius maculatus]L02
         |         |--O. chrysopsA99
         |         |--O. fulvicephalusGE05
         |         `--O. strigosusG27
              |  i. s.: ClydosmylusN91
              |--Australysmus lacustrisWHW10
                 |    |--K. falcatusA69
                 |    `--K. kimminsiWHW10
                 `--+--Porismus [Porisminae]WHW10
                    |    `--P. strigatusN91
                         |--Euporismus albatroxN91
                         |--Oedosmylus brevisWHW10
                         `--Phymatosmylus Adams 1969A69
                              `--*P. caprorum Adams 1969A69
Osmylidae incertae sedis:
  Mesosmylus atalantusP02
  Kazakhstania fasciataP02
  Epiosmylus Panfilov 1980RY02
    |--E. longicornisRY02
    `--E. panfilovi Ren & Yin 2002RY02
  Sogjuta speciosaRJ93
  ArcheosmylusRJ93 [Archeosmylidae, ArcheosmylinaeEWB18]
    `--A. complexus Whalley 1988RJ93

*Type species of generic name indicated


[A69] Adams, P. A. 1969. A new genus and species of Osmylidae (Neuroptera), from Chile and Argentina, with a discussion of planipennian genitalic homologies. Postilla 141: 1–11.

[A99] Ax, P. 1999. Das System der Metazoa II. Ein Lehrbuch der phylogenetischen Systematik. Gustav Fisher Verlag: Stuttgart (translated: 2000. Multicellular Animals: The phylogenetic system of the Metazoa vol. 2. Springer).

[EWB18] Engel, M. S., S. L. Winterton & L. C. V. Breitkreuz. 2018. Phylogeny and evolution of Neuropterida: where have wings of lace taken us? Annual Review of Entomology 63: 531–551.

[G27] Girault, A. A. 1927. Thysanoptera nova Australiensis. II. Privately published (reprinted: Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 214–215).

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[N91] New, T. R. 1991. Neuroptera (lacewings). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 525–542. Melbourne University Press: Carlton (Victoria).

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[P02] Ponomarenko, A. G. 2002. Superorder Myrmeleontidea Latreille, 1802 (=Neuropteroidea Handlirsch, 1903). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 176–189. Kluwer Academic Publishers: Dordrecht.

[RY02] Ren D. & Yin J.-C. 2002. [A new Middle Jurassic species of Epiosmylus from Inner Mongolia, China (Neuroptera: Osmylidae)]. Acta Zootaxonomica Sinica 27 (2): 274–277.

[RY03] Ren, D., & J. Yin. 2003. New ‘osmylid-like’ fossil Neuroptera from the Middle Jurassic of Inner Mongolia, China. Journal of the New York Entomological Society 111 (1): 1–11.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[R70a] Riek, E. F. 1970a. Fossil history. In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 168–186. Melbourne University Press.

[R70b] Riek, E. F. 1970b. Neuroptera (lacewings). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 472–494. Melbourne University Press.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[WHW10] Winterton, S. L., N. B. Hardy & B. M. Wiegmann. 2010. On wings of lace: phylogeny and Bayesian divergence time estimates of Neuropterida (Insecta) based on morphological and molecular data. Systematic Entomology 35: 349–378.

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