Osmylidae

Porismus strigatus, copyright Donald Hobern.

Belongs within: Euneuroptera.

The Osmylidae are a diverse group of moderate-sized to large lacewings that typically have camouflaging patterns on their wings. Larvae are long and slender with elongate, lance-like jaws and are most commonly found in moist riparian habitats. Some members of the clade Osmylomorpha have larvae living in drier locations (Engel et al. 2018).

Characters (from New 1991): Generally medium to large (fore wing length 12-30 mm or more), with wings often patterned with grey or brown and rather broad; antennae usually less than half as long as fore wing; three ocelli present, at least represented by raised tubercles on vertex; female genitalia with enlarged gonocoxite 9, and 2 spermathecae. Larvae long and fusiform, with long tapering straight jaws.

Osmylidae
|--SpilosmylomorphaEWB18
| |--GumillaA69 [GumillinaeEWB18]
| |--SpilosmylinaeRY03
| | |--Spilosmylus flavicornisRD77
| | `--Conchylosmylus triseriatusR70b
| `--ProtosmylinaeEWB18
| |--Paryphosmylus ornatusA69
| `--ProtosmylusP92
`--OsmylomorphaEWB18
| i. s.: EidoporismusN91 [EidoporisminaeEWB18]
| `--E. pulchellusN91
|--OsmylinaeRY03
| |--PlethosmylusA69
| `--Osmylus Latreille 1802WHW10, L02
| |--*O. maculatus [=Hemerobius maculatus]L02
| |--O. chrysopsA99
| |--O. fulvicephalusGE05
| `--O. strigosusG27
`--KempyninaeEWB18
| i. s.: ClydosmylusN91
|--Australysmus lacustrisWHW10
`--+--KempynusWHW10
| |--K. falcatusA69
| `--K. kimminsiWHW10
`--+--Porismus [Porisminae]WHW10
| `--P. strigatusN91
`--StenosmylinaeEWB18
|--Euporismus albatroxN91
|--Oedosmylus brevisWHW10
|--StenosmylusN91
|--CarinosmylusN91
|--StenolysmusN91
|--IsostenosmylusA69
`--Phymatosmylus Adams 1969A69
`--*P. caprorum Adams 1969A69

Osmylidae incertae sedis:
Mesosmylus atalantusP02
Kazakhstania fasciataP02
Epiosmylus Panfilov 1980RY02
|--E. longicornisRY02
`--E. panfilovi Ren & Yin 2002RY02
EuporismitesR70a
Sogjuta speciosaRJ93
KalosmylinaeR70b
ArcheosmylusRJ93 [Archeosmylidae, ArcheosmylinaeEWB18]
`--A. complexus Whalley 1988RJ93

*Type species of generic name indicated

References

[A69] Adams, P. A. 1969. A new genus and species of Osmylidae (Neuroptera), from Chile and Argentina, with a discussion of planipennian genitalic homologies. Postilla 141: 1–11.

[A99] Ax, P. 1999. Das System der Metazoa II. Ein Lehrbuch der phylogenetischen Systematik. Gustav Fisher Verlag: Stuttgart (translated: 2000. Multicellular Animals: The phylogenetic system of the Metazoa vol. 2. Springer).

[EWB18] Engel, M. S., S. L. Winterton & L. C. V. Breitkreuz. 2018. Phylogeny and evolution of Neuropterida: where have wings of lace taken us? Annual Review of Entomology 63: 531–551.

[G27] Girault, A. A. 1927. Thysanoptera nova Australiensis. II. Privately published (reprinted: Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 214–215).

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[N91] New, T. R. 1991. Neuroptera (lacewings). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 525–542. Melbourne University Press: Carlton (Victoria).

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[P02] Ponomarenko, A. G. 2002. Superorder Myrmeleontidea Latreille, 1802 (=Neuropteroidea Handlirsch, 1903). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 176–189. Kluwer Academic Publishers: Dordrecht.

[RY02] Ren D. & Yin J.-C. 2002. [A new Middle Jurassic species of Epiosmylus from Inner Mongolia, China (Neuroptera: Osmylidae)]. Acta Zootaxonomica Sinica 27 (2): 274–277.

[RY03] Ren, D., & J. Yin. 2003. New ‘osmylid-like’ fossil Neuroptera from the Middle Jurassic of Inner Mongolia, China. Journal of the New York Entomological Society 111 (1): 1–11.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[R70a] Riek, E. F. 1970a. Fossil history. In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 168–186. Melbourne University Press.

[R70b] Riek, E. F. 1970b. Neuroptera (lacewings). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 472–494. Melbourne University Press.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[WHW10] Winterton, S. L., N. B. Hardy & B. M. Wiegmann. 2010. On wings of lace: phylogeny and Bayesian divergence time estimates of Neuropterida (Insecta) based on morphological and molecular data. Systematic Entomology 35: 349–378.

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