Otitesellini

 Undescribed species of Lipothymus, photographed by Jean-Yves Rasplus.

Belongs within: Pteromalidae.

The Otitesellini are a group of wasps that oviposit on figs, either by entering the ostiole or through the fig wall. The larvae feed on the ovarium tissue, which swells to a gall (Bouček 1988).

The Sycoryctinae are a group of fig-wasps in which the females have a long, extruded ovipositor (at least partially covered by the extended last gastral tergite[s] in some Sycoryctini) and an undivided scutellum (Bouček 1988).

<==Otitesellini [Otitesellinae, Sycoecinae, Sycoecini, Sycoryctinae, Sycoryctini]BM22
    |--PhilosycusB88
    |--Eujacobsonia Grandi 1923B88
    |    `--*E. mirabilisB88
    |--Guadalia Wiebes 1867B88
    |    `--*G. vissali Wiebes 1967B88
    |--Micrognathophora Grandi 1921B88
    |    `--*M. leptopteraB88
    |--Lipothymus Grandi 1921B88
    |    `--*L. sumatranusB88
    |--Grandiana Wiebes 1961B88
    |    |--*G. wassae Wiebes 1867B88
    |    `--G. armadillo Bouček 1988B88
    |--Micranisa Walker 1875 [incl. Epicolystichus Girault 1915, Sycobiella Westwood 1883]B88
    |    |--*M. pteromaloides [=Idarnes pteromaloides]B88
    |    |--M. aereicorpus (Girault 1915) [=*Epicolystichus aereicorpus]B88
    |    `--*Sycobiella’ saundersiiB88
    |--Otitesella Westwood 1883MH11, B88 [incl. Paracolystichus Girault 1915B88]
    |    |--*O. digitataB88
    |    |--O. amplipennis (Girault 1915) [=Paracolystichus amplipennis]B88
    |    |--O. claraeB88
    |    `--O. compressiventris (Girault 1915) [=*Paracolystichus compressiventris]B88
    |--Walkerella Westwood 1883LXH13, B88 [incl. Asemantoideus Girault 1916B88, Terastiozoon Grandi 1921B88]
    |    |--*W. temeraria [incl. Terastiozoon benjamini Joseph 1957]B88
    |    |--W. dubia [=*Asemantoideus dubius]B88
    |    |--*Terastiozoon’ jacobsoniB88
    |    |--W. kurandensis Bouček 1988B88
    |    `--W. yashiroi [=Otitesella yashiroi]B88
    |--Apocrypta Coquerel 1855 (see below for synonymy)B88
    |    |--*A. perplexaB88
    |    |--A. acaeta Ulenberg in Ulenberg & Pelt 1985B88
    |    |--A. caudata (Girault 1915) [=*Goniogastrella caudatus]B88
    |    |--A. centrocampta Ulenberg in Ulenberg & Pelt 1985B88
    |    |--A. channingi (Girault 1913) [=*Idarnoides channingi]B88
    |    |--A. macroseta Ulenberg in Ulenberg & Pelt 1985B88
    |    |--A. mega Ulenberg in Ulenberg & Pelt 1985B88
    |    |--A. megischia Ulenberg in Ulenberg & Pelt 1985B88
    |    `--A. meromassa Ulenberg in Ulenberg & Pelt 1985B88
    |--Dobunabaa Bouček 1988B88
    |    `--*D. flava Bouček 1988B88
    |--Tenka Bouček 1988B88
    |    `--*T. percaudata Bouček 1988B88
    |--Watshamiella Wiebes in Bouček, Watsham & Wiebes 1981 [=Metidarnes Riek 1970 (n. n.)]B88
    |    |--*W. alataB88
    |    `--W. aurea (Girault 1919) [=Philotrypesis aurea]B88
    |--Philotrypesis Förster 1878 (see below for synonymy)B88
    |    |--*P. longicauda Förster 1878B88
    |    |--*Sycoscaptella’ affinisB88
    |    |--P. angela Girault 1915GM79
    |    |--‘Sycoscaptella’ anguliceps [=*Sycoscapterella anguliceps]B88
    |    |--P. aterrima (Saunders 1883) [=Idarnella aterrima]B88
    |    |--P. caricae [=Cynips caricae, *Idarnodes caricae]B88
    |    |--P. immaculata Girault 1915 [incl. P. longicauda Girault & Dodd in Girault 1915 non Förster 1878]B88
    |    |--*Philotrypomorpha’ indicaB88
    |    |--P. longiventris Girault 1915B88
    |    |--*Polanisa’ luteaB88
    |    |--‘Sycoscaptella’ quadrisetosa [=*Tetranemopteryx quadrisetosa]B88
    |    |--P. silvensis Girault 1915B88
    |    `--‘Idarnes’ transiens [=*Idarnella transiens]B88
    |--Sycoscapter Saunders in Westwood 1883 (see below for synonymy)B88
    |    |--*S. insignisB88
    |    |--S. australis (Froggatt 1900)B88 [=Idarnis (l. c.) australisB88, Idarnes australisG28]
    |    |--S. conocephalus Wiebes 1964B88
    |    |--S. huberi (Girault 1919) [=*Philotrypesella huberi, Sycoryctes huberi]B88
    |    |--*Sycorycteridea’ keralensisB88
    |    |--S. miltoni (Girault 1931) [=*Ideoidarnes miltoni]B88
    |    |--‘Sycoscapter’ monilifer [=*Sycoscapteridea monilifer]B88
    |    |--*Sycoryctes’ patellarisB88
    |    |--*Arachonia’ plumosaB88
    |    |--S. subaeneus (Girault 1915) [=*Idarnomorpha subaenea, Idarnes subaeneus]B88
    |    `--S. varicilia (Girault 1928) [=Sycoryctes varicilia]B88
    |--CrossogasterB88
    |--Sycoecus Waterston 1914BM22
    |--Seres wardiBM22
    |--Diaziella Grandi 1928B88
    |    |--*D. bicolorB88
    |    `--D. falcataB88
    |--Robertsia Bouček 1988B88
    |    |--*R. mandibularis Bouček 1988B88
    |    `--R. xylosyciae Bouček 1988B88
    |--MarginaliaBM22
    |--AdiyodiellaBM22
    |--BoucekaBM22
    |--ComptoniellaBM22
    |--Grasseiana Abdurahiman & Joseph 1967BM22, B88
    |    |--*G. callosaB88
    |    `--G. boschmai (Wiebes 1964) [=Sycobiella boschmai]B88
    |--PhilosycellaBM22
    `--PhiloverdanceBM22

Apocrypta Coquerel 1855 [incl. Agaonella Baker 1913, Goniogaster Mayr 1885, Goniogastrella Girault & Dodd in Girault 1915, Idarnoides Girault 1913; Apocryptini]B88

Philotrypesis Förster 1878 [incl. Idarnella Westwood 1883, Idarnodes Westwood 1883, Philotrypomorpha Abdurahiman & Joseph 1976, Polanisa Walker 1875 (nom. rej.), Sycoscaptella Westwood 1883, Sycoscapterella Ashmead 1904, Tetranemopteryx Ashmead 1904; Philotrypesini]B88

Sycoscapter Saunders in Westwood 1883 [incl. Arachonia Joseph 1957, Idarnomorpha Girault 1915, Ideoidarnes Girault 1931, Philotrypesella Girault 1919, Sycorycteridea Abdurahiman & Joseph 1975, Sycoryctes Mayr 1885, Sycoscapteridea Ashmead 1904]B88

*Type species of generic name indicated

References

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[BM22] Burks, R., M.-D. Mitroiu, L. Fusu, J. M. Heraty, P. Janšta, S. Heydon, N. D.-S. Papilloud, R. S. Peters, E. V. Tselikh, J. B. Woolley, S. van Noort, H. Baur, A. Cruaud, C. Darling, M. Haas, P. Hanson, L. Krogmann & J.-Y. Rasplus. 2022. From hell’s heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera). Journal of Hymenoptera Research 94: 13–88.

[G28] Girault, A. A. 1928. A prodigeous discourse on wild animals. Privately published (reprinted: Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 218–220).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1–400.

[LXH13] Li, Z., H. Xiao & D.-W. Huang. 2013. Sirovena Bouček (Pteromalidae: Pireninae), a new member of the fig wasp community associated with Ficus microcarpa (Moraceae). Zootaxa 3619 (5): 581–588.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PloS One 6 (11): e27023.

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