Palaeocopida

Left valve of Kirkbyella delicata, from Hoare & Merrill (2004).

Belongs within: Ostracoda.
Contains: Youngielloidea, Paraparchitidae, Kirkbyelloidea, Glyptopleuroidea, Tetradellidae, Sigmoopsidae, Beyrichioidea, Eurychilinidae, Oepikellidae, Hollinidae, Piretellidae, Ctenonotellidae, Hollinella, Tvaerenellidae, Aparchitidae, Punciocopida, Aechminellidae, Bollioidea, Aechminidae, Drepanellidae.

The Drepanellacea are a group of non-dimorphic ostracods known from the Ordovician to the Permian (Whatley et al. 1993). Members of the group vary greatly in development of nodes and features of the marginal rim but are united by the existence of intermediate forms (Benson et al. 1961).

<==Palaeocopida [Beyrichiida, Beyrichiocopa, Leiocopa, Paleocopa, Palaeocopina]L05
|--Beyrichiina [Hollinomorphinina]L05
| |--TetradelloideaL05
| | |--TetradellidaeBB61
| | |--SigmoopsidaeL05
| | `--Ctenoloculina Bassler 1941BB61 [CtenoloculinidaeL05, Ctenoloculininae]
| | `--*C. cicatricosa (Warthin 1934) [=Tetradella cicatricosa]BB61
| `--CavicarinataL05
| |--BeyrichioideaL05
| |--Eurychilinoidea [Eurychilinacea, Hollinacea, Hollinoidea, Oepikellacea]L05
| | | i. s.: Echinoprimitia Harris 1957BB61
| | | `--*E. imputata Harris 1957BB61
| | |--EurychilinidaeTM02
| | |--OepikellidaeTM02
| | |--PribylitidaeBB61
| | | |--Pribylites Pokorný 1950 [incl. Parapribylites Pokorný 1950]BB61
| | | | `--*P. moravicus Pokorný 1950BB61
| | | |--Mirochilina Bouček 1936BB61
| | | | `--*M. jarovensis Bouček 1936BB61
| | | `--Sphenicibysis Kesling 1952 [=Sphenicibys (l. c.)]BB61
| | | `--*S. hypoderota Kesling 1952BB61
| | |--HollinidaeL05
| | |--PiretellidaeBB61
| | |--CtenonotellidaeWSB93
| | |--HollinellaWSB93
| | |--SoanellidaeWSB93
| | | |--Fuscinullina pectinata Kanygin 1967WSB93
| | | `--Sibiretella furcata Kanygin 1967WSB93
| | |--Oepikium Agnew 1942BB61 [incl. Biflabellum Öpik 1935 non Döderlein 1913BB61; OepikiidaeTM02, Oepikiumidae]
| | | |--*O. tenerum (Öpik 1935) [=Biflabellum tenerum]BB61
| | | |--O. novum Sarv 1959WSB93
| | | `--O. porkuniensis Henningsmoen 1954WSB93
| | |--Egorovella Ivanova 1959LH61 [EgorovellidaeWSB93]
| | | |--*E. compacta Ivanova 1959LH61
| | | |--E. defecta Ivanova 1959WSB93
| | | `--E. poricostata Kanygin 1965WSB93
| | |--SarvinidaeWSB93
| | | |--Anticostiella Copeland 1973B94
| | | | `--*A. ellisensis Copeland 1973B94
| | | `--Distobolbina Sarv 1959BB61
| | | |--*D. nabalaensis Sarv 1959BB61
| | | `--D. pinna Schallreuter 1964WSB93
| | |--DolborellidaeWSB93
| | | |--Planiprimites solitus Kanygin 1967WSB93
| | | |--Costaegera hastaa Abushik 1960WSB93
| | | `--Dolborella Ivanova 1959LH61
| | | `--*D. plana Ivanova 1959LH61
| | |--Chilobolbinidae [Chilobolbininae]BB61
| | | |--Cystomatochilina Jaanusson 1957BB61
| | | | `--*C. umbonata (Krause 1892) [=Primitia (Ulrichia) umbonata]BB61
| | | `--Chilobolbina Ulrich & Bassler 1923 [incl. Chilobolba Bonnema 1938]BB61
| | | |--*C. dentifera (Bonnema 1909) [=Primitia dentifera]BB61
| | | `--C. hartfordensisBB61
| | |--Euprimitiidae [Euprimitiinae]WSB93
| | | |--Steinfurtia macroreticulata (Hessland 1949)WSB93
| | | |--Gryphiswaldensia visbya Schrallreuter 1969WSB93
| | | `--Euprimitia Ulrich & Bassler 1923BB61
| | | |--*E. sanctipauli (Ulrich 1894) [=Primitia sanctipauli]BB61
| | | `--E. labiosaBB61
| | `--TvaerenellidaeBB61
| |--Aparchitoidea [Aparchitacea, Leiocopinina]L05
| | |--AparchitidaeHM04
| | |--SchmidtellidaeWSB93
| | | |--Punctoschmidtella umbopunctata (Harris 1957)WSB93
| | | |--Schmidtella Ulrich 1892BB61
| | | | |--*S. crassimarginata Ulrich 1892BB61
| | | | `--S. umbonataBB61
| | | `--Paraschmidtella Swartz 1936WSB93, BB61
| | | |--*P. dorsopunctata Swartz 1936BB61
| | | `--P. ovataBB61
| | `--GlanditidaeL05
| `--PunciocopidaL05
|--Binodicopina [Aechminacea, Drepanellacea]L05
| | i. s.: SpinigeritidaeWSB93
| | |--Susus sus Schallreuter 1991WSB93
| | `--Eocytherella Bonnema 1933WSB93, BB61
| | |--E. nioni Vannier 1986WSB93
| | |--E. smithi (Jones 1887) [=Cytherella smithi]BB61
| | `--E. troedssoni (Bonnema 1933) (see below for synonymy)BB61
| | AechminellidaeBB61
| |--BollioideaL05
| |--AechminidaeL05
| |--DrepanellidaeL05
| |--Nodella Zaspelova 1952BB61 [Nodellacea, Nodellidae, Nodellinae, NodelloideaL05]
| | |--*N. svinordensis Zaspelova 1952BB61
| | |--N. faceata Rozhdestvenskaya 1972WSB93
| | `--N. lefrevei Becker 1971WSB93
| `--LimbatuloideaL05
| |--Neodrepanella Zaspelova 1952BB61 (see below for synonymy)
| | `--*N. tricornis (Batalina 1941) [=Drepanella tricornis]BB61
| `--Ulrichia Jones 1890BB61 [incl. Limbatula Zaspelova 1952BB61, Warthinia Spivey 1939BB61; LimbatulidaeL05]
| |--*U. conradi Jones 1890BB61
| `--U. nodosa [=*Warthinia nodosa]BB61
`--Eridostracina [Eridostraca, Eridostracoda]BT04
|--Conchoprimitia Öpik 1935BB61 (see below for synonymy)
| |--*C. gammae Öpik 1935BB61
| |--C. symmetricaBB61
| `--C. tolli (Bonnema 1909)WSB93
|--Eridoconcha Ulrich & Bassler 1923BB61 [EridoconchidaeWSB93, Eridoconchinae]
| |--*E. rugosa Ulrich & Bassler 1923BB61
| |--E. marginataBB61
| |--E. multiannulataBB61
| `--E. simpsoni Harris 1931WSB93
|--Cryptophyllus Levinson 1951HM04 [CryptophyllidaeL05]
| |--*C. oboloides (Ulrich & Bassler 1923) [=Eridoconcha oboloides]BB61
| |--C. magnus (Harris 1931)WSB93
| `--C. sulcatusBB61
`--SchaefericonchidaeL05

Palaeocopida incertae sedis:
Kozlowskiella Přibyl 1953BB61
`--*K. kozlowskii (Přibyl 1953) [=Ulrichia (*Kozlowskiella) kozlowskii]BB61
Acanthobolbina Harris 1957BB61
`--*A. loeblichi Harris 1957BB61
Dicranella Ulrich 1894 non (Müll.Hal.) Schimp. 1856 (ICBN)BB61
|--*D. bicornis Ulrich 1894BB61
`--D. marginataBB61
Haplobolbina Harris 1957BB61
`--*H. arcuata Harris 1957BB61
Karlsteinella Bouček 1936BB61
`--*K. reticulata Bouček 1936BB61
Nanopsis Henningsmoen 1954BB61
`--*N. nanella (Moberg & Segerberg 1906) [=Beyrichia nanella]BB61
JudahellaJB12
RetisacculusJB12
Eopilla ingelorae Schallreuter 1993BT04
YoungielloideaBB61
Tibolbinidae [Tribolbinacea]WSB93
|--Kolmodinia grandis (Kolmodin 1869)WSB93
`--Tribolbina Latham 1932BB61
|--*T. carnegiei Latham 1932BB61
`--T. doescheri Sohn 1978WSB93
Paraparchitoidea [Paraparchitacea, Paraparchitocopina]L05
|--ParaparchitidaeHM04
`--JaanussonidaeWSB93
|--Jaanussonia unicerata Schallreuter 1971WSB93
`--Kayina Harris 1957WSB93, BB61
`--*K. hybosa Harris 1957BB61
KirkbyelloideaL05
GlyptopleuroideaL05
Ectoprimitiodes Berdan 1988B94
`--E. moorei (Harris 1957) (see below for synonymy)B94

Conchoprimitia Öpik 1935BB61 [incl. Conchoides Hessland 1949BB61, Hyperchilarina Harris 1957BB61; ConchoprimitiidaeWSB93, Conchoprimitiinae]

Ectoprimitiodes moorei (Harris 1957) [=Eoprimitia moorei; incl. Eo. dimunicarina Kraft 1962, *Ectoprimitiodes dimunicarina]B94

Eocytherella troedssoni (Bonnema 1933) [=Cytherella troedssoni, Primitia tenera Troedsson 1918 (preoc.)]BB61

Neodrepanella Zaspelova 1952BB61 [incl. Tetracornella Zaspelova 1952BB61; NeodrepanellidaeL05, Neodrepanellinae]

*Type species of generic name indicated

References

[B94] Becker, G. 1994. A remarkable Ordovician ostracod fauna from Orphan Knoll, Labrador Sea. Scripta Geologica 107: 1–25.

[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[BT04] Braddy, S. J., V. P. Tollerton Jr, P. R. Racheboeuf & R. Schallreuter. 2004. Eurypterids, phyllocarids, and ostracodes. In: Webby, B. D., F. Paris, M. L. Droser & I. G. Percival (eds) The Great Ordovician Biodiversification Event pp. 255–265. Columbia University Press.

[HM04] Hoare, R. D., & G. K. Merrill. 2004. A Pennsylvanian (Morrowan) ostracode fauna from Texas. Journal of Paleontology 78 (1): 185–204.

[JB12] Johnson, M. E., & B. G. Baarli. 2012. Development of intertidal biotas through Phanerozoic time. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 63–128. Springer.

[LH61] Levinson, S. A., & H. V. Howe. 1961. Addendum. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q421–Q422. Geological Society of America and University of Kansas Press.

[L05] Liebau, A. 2005. A revised classification of the higher taxa of the Ostracoda (Crustacea). Hydrobiologia 538: 115–137.

[TM02] Tinn, O., & T. Meidla. 2002. An enigmatic early ?palaeocope ostracode from the Arenig of NW Russia. Acta Palaeontologica Polonica 47: 685–690.

[WSB93] Whatley, R. C., D. J. Siveter & I. D. Boomer. 1993. Arthropoda (Crustacea: Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 343–356. Chapman & Hall: London.

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