Pholcinae

Pholcus phalangioides, copyright Olaf Leillinger.
Belongs within: Pholcidae.

The Pholcinae are a group of daddy-long-legs spiders united by the reduction of the tarsal comb hairs to a single ventral row, and the usual presence on the male chelicerae of a pair of proximal lateral apophyses (Huber 2011).

<==Pholcinae
|  i. s.: Aetana Huber 2005 H11
|         Wanniyala Huber & Benjamin 2005 H11
|–+–Nyikoa Huber 2007 H11
|  |    `–N. limbe Huber 2007 HF08
|  `–Zatavua Huber 2003 H11
|       |–Z. vohiparara Huber 2003 HF08
|       `–Z. zanahary H11
`–+–Paraspermophora Wunderlich 2004 H11
|–Anansus Huber 2007 H11
|–Savarna Huber 2005 H11
|–Khorata Huber 2005 H11
|    `–K. khammouan Huber 2005 HF08
|–+–+–Paramicromerys Millot 1946 H11
|  |  |    `–P. scharffi Huber 2003 HF08
|  |  `–+–Buitinga Huber 2003 HF08, H11
|  |     |    |–B. asax Huber 2003 HF08
|  |     |    `–B. kadogo H11
|  |     `–Spermophorides Wunderlich 1992 HF08, H11
|  |          |–S. cuneata (Wunderlich 1987) HF08
|  |          `–S. elevata (Simon 1873) HF08
|  `–+–Metagonia Simon 1893 H11
|     |    |–M. conica (Simon 1893) HF08
|     |    |–M. mariguitarensis (González-Sponga 1998) T09
|     |    `–M. reventazona Huber 1997 HF08
|     `–Spermophora Hentz 1841 HF08, H11
|          |–S. kerinci Huber 2005 HF08
|          |–S. senoculata (Dugès 1836) H11
|          `–S. usambara Huber 2003 HF08
`–+–Quamtana Huber 2003 H11
|    `–Q. bonamanzi Huber 2003 HF08
|–Belisana Thorell 1898 H11
|    |–B. airai Huber 2005 HF08
|    |–B. fiji Huber 2005 HF08
|    |–B. hormigai H11
|    `–B. ketambe H11
`–+–Micropholcus Deeleman-Reinhold & Prinsen 1987 H11
|    `–M. fauroti (Simon 1887) H11
|–Pehrforsskalia Deeleman-Reinhold & van Harten 2001 H11
|    `–P. conopyga Deeleman-Reinhold & van Harten 2001 HF08
`–+–+–Panjange Deeleman-Reinhold & Deeleman 1983 HF08, H11
|  |    `–P. mirabilis Deeleman-Reinhold 1986 HF08
|  `–Leptopholceae H11
|       |–Micromerys Bradley 1877 H11
|       |    `–M. daviesae Deeleman-Reinhold 1986 HF08
|       `–Leptopholcus Simon 1893 H11
|            |–L. debakkeri H11
|            |–L. gracilis H11
|            `–L. hispaniola Huber 2000 HF08
`–+–Calapnita Simon 1892 HF08, H11
|    |–C. phyllicola Deeleman-Reinhold 1986 HF08
|    `–C. saluang H11
|–Ossinissa Dimitrov & Ribera 2005 HF08, H11
|    `–*O. justoi (Wunderlich 1991) [=Spermophorides justoi] F05
|–Uthina Simon 1893 HF08, H11
|    `–U. luzonica Simon 1893 HF08
`–Pholcus Walckenaer 1805 H11
|–P. barbarus Lucas 1846 E12
|–P. bourgini H11
|–P. caudatus V09
|–P. chattoni H11
|–P. crypticolens Bösenberg & Strand 1906 RA08
|–P. kwamgumi H11
|–P. lamperti H11
|–P. manueli Gertsch 1937 [incl. P. affinis Schenkel 1953] RA08
|–P. opilionoides (Schrank 1781) HF08
|–P. phalangioides (Fuesslin 1775) H11
|–P. quadripunctatus Lucas 1846 E12
|–P. quinquenotatus H11
|–P. sphaeroides B96
`–P. zichyi Kulczyński 1901 K01

*Type species of generic name indicated

REFERENCES

[B96] Brunet, B. 1996. Spiderwatch: A Guide to Australian Spiders. Reed New Holland: Sydney.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[H11] Huber, B. A. 2011. Phylogeny and classification of Pholcidae (Araneae): an update. Journal of Arachnology 39 (2): 211–222.

[HF08] Huber, B. A., & N. Fleckenstein. 2008. Comb-hairs on the fourth tarsi in pholcid spiders. Journal of Arachnology 36 (2): 232–240.

[K01] Kulczyński, V. 1901. Arachnoideák [Arachnoidea]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 311–369. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[RA08] Ramalho, M. O., D. Araujo, M. C. Schneider, A. D. Brescovit & D. M. Cella. 2008. Mesabolivar brasiliensis (Moenkhaus 1898) and Mesabolivar cyaneotaeniatus (Keyserling 1891) (Araneomorphae, Pholcidae): close relationship reinforced by cytogenetic analysis. Journal of Arachnology 36 (2): 453–456.

[T09] Taylor, C. K. 2009. Australiscutum, a new genus of Monoscutidae (Arachnida: Opiliones) from eastern Australia, with the first record of asymmetrical chelicerae in Opiliones. Insect Systematics and Evolution 40: 319–332.

[V09] Verdcourt, B. (ed.) 2009. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

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