Phylloceras sp., copyright Hectonichus.

Belongs within: Neoammonoidea.

The Phylloceratidae are a group of ammonites with mostly smooth or feebly ornamented conchs, and phylloid saddle endings of the suture line (Engeser & Keupp 2002). They have a long stratigraphic range from the early Jurassic to the end of the Cretaceous.

|--Phyllopachyceras Spath 1925 [Phyllopachyceratinae]DJ95
| |--*P. infundibulum (d’Orbigny 1841) (see below for synonymy)DJ95
| |--P. baborense (Coquand 1880) [=Ammonites baborensis, Partschiceras baborense, Phylloceras baborense]DJ95
| |--P. bontshevi Manolov 1962DJ95
| |--P. bulgaricum Dimitrova 1967DJ95
| |--P. chitinanum Imlay 1960DJ95
| |--P. crassum Drushchic 1956DJ95
| |--P. ectocostatum (Drushchic 1956)DJ95
| |--P. eichwaldiDJ95
| | |--P. e. eichwaldiDJ95
| | `--P. e. occidentale (Wiedmann 1964) [=Partschiceras eichwaldi occidentale]DJ95
| |--P. katschiense Drushchic 1956DJ95
| |--P. ladinumDJ95
| |--P. oregonense (Anderson 1938) (see below for synonymy)DJ95
| |--P. rouyanumDJ95
| |--P. segre Drushchic 1956DJ95
| `--P. vergonsense Delanoy & Joly 1995DJ95
|--Calliphylloceratinae [Holcophylloceratinae, Ptychophylloceratinae]DJ95
| |--Holcophylloceras Spath 1927 [incl. Salfeldiella Spath 1927, Telegdiceras Kovacs 1942]WCH96
| | |--*H. mediterraneum (Neumayr 1871) [=Phylloceras mediterraneum]WCH96
| | `--H. calypso [=Sowerbyceras (Holcophylloceras) calypso]WK81
| |--Sowerbyceras Parona & Bonarelli 1894WCH96, EK02 (see below for synonymy)
| | |--*S. tortisulcatus (Orbigny 1841) [=Ammonites tortisulcatus]WCH96
| | `--S. loryiZ81
| |--Neocalliphylloceras Besairie 1936DJ95
| | |--*N. collignoni Besairie 1936DJ95
| | `--N. sablyense (Karakasch 1907)DJ95
| `--Calliphylloceras Spath 1927 [incl. Capitanioceras Kovacs 1939]WCH96
| | i. s.: C. freibrockiP87
| | C. heterophylloidesP87
| | C. psilomorphumP93
| |--*C. (Calliphylloceras) disputabile (Zittel 1869) [=Phylloceras disputabile]WCH96
| `--C. (Ptychophylloceras Spath 1927)EK02 (see below for synonymy)
| |--*Ptychophylloceras’ feddeni (Waagen 1875) [=Phylloceras feddeni]WCH96
| `--C. (P.) ptychoideum Spath 1927EK02
`--Phylloceratinae [Pseudophylloceratinae]DJ95
|--Adabofoloceras Joly 1975P87
|--Hyporbulites Breistroffer 1947DJ95
| `--*H. seresitensis (Pervinquière 1907) [=Phylloceras velledae var. seresitensis]DJ95
|--Carinophylloceras Klinger, Wiedmann & Kennedy 1975WCH96
| `--*C. collignoni Klinger, Wiedmann & Kennedy 1975WCH96
`--Phylloceras Suess 1865DJ95
| i. s.: P. billingsi (Meek 1859) [=Ammonites billingsi]EK02
| P. kudernatschiEK02
| P. kunthi Neumayr 1971EK02
| P. onoenseWK81
| P. spathiK79
| P. subatlas Voronetz 1962EK02
|--P. (Phylloceras)DJ95 (see below for synonymy)
| `--*P. (P.) heterophyllum (Sowerby 1820)DJ95 (see below for synonymy)
|--P. (Hypophylloceras Salfeld 1924)DJ95 (see below for synonymy)
| |--P. (*H.) ononense Stanton 1895DJ95
| |--P. (H.) moussoni (Ooster 1860) [=Ammonites moussoni]DJ95
| |--P. (H.) ponticuli (Rousseau 1842) (see below for synonymy)DJ95
| | |--P. p. ponticuliDJ95
| | `--P. p. plicatum Delanoy & Joly 1995DJ95
| |--‘Hypophylloceras’ subramosumTLY03
| |--P. (H.) thetysDJ95
| `--P. (H.) velledae (Michelin 1838)KK08
|--P. (Macrophylloceras Spath 1927)EK02, WCH96
| `--P. (M.) grossicostatumEK02
|--P. (Neophylloceras Spath 1927)EK02 (see below for synonymy)
| |--‘Hyphophylloceras’ (*N.) ramosus (Meek 1857) [=Ammonites (Scaphites) ramosus]WCH96
| |--P. (N.) suryaBRW98
| `--‘Neophylloceras’ velledaeformeP93
|--P. (Partschiceras Fucini 1923) (see below for synonymy)WCH96
| |--P. (*P.) ‘partschi’ (Hauer 1854) [=Ammonites partschi non Klipstein 1843]WCH96
| |--P. (P.) grantziEK02
| |--P. (P.) monestieriP87
| `--P. (P.) pacificum Frebold & Tipper 1975EK02
`--P. (Zetoceras Kovacs 1939) [incl. Lavizzaroceras Kovacs 1939]WCH96
|--P. (*Z.) zetes (Orbigny 1850) [=Ammonites zetes]WCH96
`--‘Zetoceras’ thorsteinssoni Frebold 1961P87

Calliphylloceras (Ptychophylloceras Spath 1927)EK02 [incl. Neumayriceras Sorrentino 1942 non Rollier 1909WCH96, Tatroceras Kovac 1939WCH96, Tatrophylloceras Beznosov 1957WCH96]

Phylloceras (Hypophylloceras Salfeld 1924)DJ95 [incl. Aphroditiceras Mahmoud 1955WCH96, Euphylloceras Drushchits 1953WCH96, Goretophylloceras Collignon 1948WCH96]

Phylloceras (Hypophylloceras) ponticuli (Rousseau 1842) [=Ammonites ponticuli, Calliphylloceras ponticuli, Euphylloceras ponticuli]DJ95

Phylloceras (Neophylloceras Spath 1927)EK02 [incl. Epiphylloceras Collignon 1956WCH96, Paraphylloceras Shimizu 1935 non Salfeld 1919WCH96]

Phylloceras (Partschiceras Fucini 1923) [incl. Adobofoloceras Joly 1977, Partschiphylloceras Roman 1938, Procliviceras Fucini 1923, Proclivioceras Roman 1938]WCH96

Phylloceras Suess 1865 (Phylloceras)DJ95 [incl. Heterophylloceras Kovacs 1939WCH96, Pseudophylloceras Beznosov 1957WCH96, Rhacoceras Hyatt 1867WCH96, Xeinophylloceras Buckman 1921WCH96]

*Phylloceras (Phylloceras) heterophyllum (Sowerby 1820)DJ95 [=Ammonites heterophyllusDJ95; incl. A. heterophyllus amalthei Quenstedt 1845CDH04, A. heterophyllus jurensis Quenstedt 1845CDH04, A. heterophyllus numismalis Quenstedt 1845CDH04, A. heterophyllus ornati Quenstedt 1845CDH04, A. heterophyllus posidoniae Quenstedt 1845CDH04]

*Phyllopachyceras infundibulum (d’Orbigny 1841) [=Ammonites infundibulum, Partschiceras infundibulum, Phylloceras infundibulum]DJ95

Phyllopachyceras oregonense (Anderson 1938) [=Phylloceras oregonense; incl. Phyllopachyceras myrtlense, Phyllopachyceras umquanum]DJ95

Sowerbyceras Parona & Bonarelli 1894WCH96, EK02 [incl. Gyrophyllites Wiedmann 1963 non Glocker 1841WCH96, Martelliceras Sorrentino 1942 non Schindewolf 1925WCH96]

*Type species of generic name indicated


[BRW98] Beesley, P. L., G. J. B. Ross & A. Wells (eds) 1998. Fauna of Australia vol. 5. Mollusca: The Southern Synthesis. Part A. Australian Biological Resources Study: Canberra.

[CDH04] Callomon, J. H., D. T. Donovan & M. K. Howarth. 2004. F. A. Quenstedt’s trinomial nomenclature of Jurassic ammonites. Palaeontology 47 (4): 1063–1073.

[DJ95] Delanoy, G., & B. Joly. 1995. Sur quelques Phylloceratidae (Phylloceratina, Ammonoidea) de Barrémien supérieur et de l’Aptien inférieur du sud-est de la France. Mésogée 54: 121–138.

[EK02] Engeser, T., & H. Keupp. 2002. Phylogeny of the aptychi-possessing Neoammonoidea (Aptychophora nov., Cephalopoda). Lethaia 35: 79–96.

[K79] Kauffman, E. G. 1979. Cretaceous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A418–A487. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[KK08] Kennedy, W. J., & H. C. Klinger. 2008. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Lyelliceratinae Spath, 1921. African Natural History 4: 57–111.

[P93] Page, K. N. 1993. Mollusca: Cephalopoda (Ammonoidea: Phylloceratina, Lytoceratina, Ammonitina and Ancyloceratina). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 213–227. Chapman & Hall: London.

[P87] Poulton, T. P. 1987. Zonation and correlation of Middle Boreal Bathonian to Lower Callovian (Jurassic) ammonites, Salmon Cache Canyon, Porcupine River, northern Yukon. Geological Survey of Canada—Bulletin 358: 1–155.

[TLY03] Tanabe, K., N. H. Landman & Y. Yoshioka. 2003. Intra- and interspecific variation in the early internal shell features of some Cretaceous ammonoids. Journal of Paleontology 77 (5): 876–887.

[WK81] Wiedmann, J., & J. Kullmann. 1981. Ammonoid sutures in ontogeny and phylogeny. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 215–255. Academic Press.

[WCH96] Wright, C. W., J. H. Calloman & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[Z81] Ziegler, B. 1981. Ammonoid biostratigraphy and provincialism: Jurassic-Old World. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 433–457. Academic Press.

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