Pink mountain-heath Phyllodoce empetriformis, copyright Walter Siegmund.

Belongs within: Ericaceae.

The Phyllodoceae are a clade of boreal shrubs with flowers on articulated pedicels.

Characters (from Kron et al. 2002): Shrubs. Leaves evergreen, alternate and spiral, decussate, or whorled, ericoid or non-ericoid, entire to serrulate, convolute or revolute in bud. Leaf epidermal cells not lignified. Indumentum of glandular hairs and elongate non-glandular hairs, and sometimes papillae. Inflorescence terminal, rarely axillary, spicate, racemose, or corymbose, sometimes reduced; bracts paired and more or less basal, bracteoles usually present; calyx articulated with pedicel. Flowers 4–6-merous, actinomorphic. Calyx lobes variable in size, withering persistent or persistent; corolla sympetalous or choripetalous, rotate to urceolate, campanulate, the lobes small to large. Stamens 5–10, included or exserted, the filaments straight to curved, unicellular-pubescent or smooth; anthers smooth or roughened, lacking appendages, dehiscing by slits or terminal pores. Pollen usually with viscin threads. Ovary 2–6-locular, with axile placentation; style impressed or not; stigma truncate, sometimes slightly expanded. Fruit a septicidal capsule, sometimes partly loculicidal as well; seeds with testa cells more or less isodiametric to elongated; embryo with two cotyledons.

<==Phyllodoceae [Cladothamneae, Epigaeae, Leiophylleae, Phyllodocoideae]
    |    |--E. bracteataKJ02
    |    `--E. racemosaKJ02
    `--+--KalmiaKJ02 [incl. DendriumBGS05, LoiseleuriaKJ02]
       |    |--K. angustifoliaKJ02
       |    |--K. polifoliaKJ02
       |    |    |--K. p. ssp. polifoliaH93
       |    |    `--K. p. ssp. microphyllaH93
       |    `--+--K. latifoliaKJ02
       |       `--+--K. buxifolia (Bergius) Gift, Kron & Stevens in Kron, Judd et al. 2002KJ02
       |          `--K. procumbens (Linnaeus) Gift, Kron & Stevens in Kron, Judd et al. 2002 (see below for synonymy)KJ02
       `--+--+--Epigaea repensKJ02
          |  `--Rhodothamnus chamaecistusKJ02
          `--+--Kalmiopsis leachianaKJ02
                  |--P. breweriW81
                  |--P. caeruleaKJ02
                  |--P. empetriformisKJ02
                  |--P. nipponicaKJ02
                  `--P. taxifolia [incl. Menziesia coerulea]C55

Kalmia procumbens (Linnaeus) Gift, Kron & Stevens in Kron, Judd et al. 2002 [=Azalea procumbens, Loiseleuria procumbens]KJ02

*Type species of generic name indicated


[BGS05] Banfi, E., G. Galasso & A. Soldano. 2005. Notes on systematics and taxonomy for the Italian vascular flora. 1. Atti Soc. It. Sci. Nat. Museo Civ. Stor. Nat. Milano 146 (2): 219–244.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[KJ02] Kron, K. A., W. S. Judd, P. F. Stevens, D. M. Crayn, A. A. Anderberg, P. A. Gadek, C. J. Quinn & J. L. Luteyn. 2002. Phylogenetic classification of Ericaceae: molecular and morphological evidence. Botanical Review 68: 335–423.

[W81] West, E. W. 1981. Adaptive patterns in behavior of the Sierran pika, Ochotona princeps. Dissertation Abstracts International B 41: 4374.

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