Phyllotocus

 Phyllotocus macleayi caught in a spider web, copyright Mike Gordon.

Belongs within: Melolonthinae.

Phyllotocus is an Australian genus of often relatively brightly coloured chafer beetles. The antennae are nine-segmented in species of the P. macleayi species group but eight-segmented in other species (Britton 1957).

Characters (from Britton 1957): Labrum transverse, clearly separated from the clypeus by a suture; clypeus reflexed at sides, without a transverse ridge, the general surface of clypeus and frons slightly convex; antennae eight- or nine-segmented, including a three-segmented club; tarsal claws slightly curved, slender, simple, inner claw of anterior tarsus usually enlarged in male; posterior angle of pronotum obtuse or rounded, not closely applied to shoulders of the elytra; maxillae with lacinia rounded and setose; prosternum with a long slender process in the middle just behind anterior coxae, process with strong spines at apex.

<==Phyllotocus Fischer 1823 [incl. Macrothops Boisduval 1835, Macrotops (l. c.)]HW92
    |  i. s.: P. lottini (Boisduval 1835)M86, HW92 [=Macrothops lottiniiHW92]
    |         P. marginalis Bohem. 1858M86
    |--P. macleayi groupB57
    |    |--*P. macleayi Fischer von Waldheim 1823HW92, A06
    |    |    |--P. m. macleayiHW92 (see below for synonymy)
    |    |    `--P. m. pallidus Lea 1919HW92
    |    |--P. bimaculatus Erichson 1842B57, HW92
    |    `--P. nigripennis Lea 1919B57, HW92 [=P. bimaculatus nigripennisHW92]
    |--P. meyricki groupB57
    |    |--P. chapmani Britton 1957B57, HW92
    |    |--P. darlingtoni Britton 1957B57, HW92
    |    |--P. gilvus Britton 1957B57, HW92
    |    |--P. laterofuscus Lea 1919B57, HW92
    |    |--P. luridus Britton 1957 non Cheiragra lurida Macleay 1864 (not preoc. if in dif. gen.)B57, HW92
    |    |--P. melanurus Britton 1957B57, HW92
    |    |--P. meyricki Blackburn 1888HW92
    |    |--P. occidentalis Blackburn 1888B57, HW92 [incl. P. occidentalis apicifuscus Lea 1919HW92]
    |    `--P. pelus Britton 1957B57, HW92
    |--P. rufipennis groupB57
    |    |--P. apicalis Macleay 1864B57, HW92 [incl. P. navicularis erythroderes Lea 1919HW92]
    |    |--P. armstrongi Britton 1957B57, HW92
    |    |--P. australis (Boisduval 1835)B57, HW92 (see below for synonymy)
    |    |--P. erythropterus Blanchard 1850B57, HW92
    |    |--P. giveni Britton 1957B57, HW92
    |    |--P. kingii Macleay 1864B57, HW92
    |    |--P. marginipennis Macleay 1864B57, HW92 [incl. P. marginatus Macleay 1864HW92]
    |    |--P. navicularis Blanchard 1850B57, HW92 (see below for synonymy)
    |    |--P. ruficollis Macleay 1864B57, HW92 [incl. P. sericeus Macleay 1871HW92]
    |    |--P. rufipennis (Boisduval 1835) [=Macrothops rufipennis, Macrotops rufipennis]B57
    |    `--P. variicollis Macleay 1871B57, HW92
    |--P. scutellaris groupB57
    |    |--P. moestus (Boisduval 1835)B57, HW92 (see below for synonymy)
    |    `--P. scutellaris Macleay 1864HW92
    `--P. ustulatus groupB57
         |--P. basalis Lea 1919B57, HW92 [=P. bimaculatus basalisHW92; incl. P. bimaculatus insularis Lea 1919HW92]
         |--P. decipiens Lea 1919B57, HW92
         `--P. ustulatus Blanchard 1850HW92

Phyllotocus australis (Boisduval 1835)B57, HW92 [=Macrothops australisHW92; incl. P. discoidalis Burmeister 1855HW92, P. iridescens Macleay 1864HW92, P. palliatus Macleay 1864HW92]

*Phyllotocus macleayi macleayi Fischer 1823HW92 [incl. P. assimilis Macleay 1864HW92, Macrothops praeusta Boisduval 1835HW92, P. praeustusM86]

Phyllotocus moestus (Boisduval 1835)B57, HW92 [=Macrothops moestaHW92; incl. P. marginicollis Boheman 1858HW92, P. oblongus Boheman 1858HW92, P. velutinus Boheman 1858HW92]

Phyllotocus navicularis Blanchard 1850B57, HW92 [incl. P. navicularis apicalis Lea 1919 non Macleay 1864HW92, P. navicularis rufibasis Lea 1919HW92]

*Type species of generic name indicated

References

[A06] Ahrens, D. 2006. The phylogeny of Sericini and their position within the Scarabaeidae based on morphological characters (Coleoptera: Scarabaeidae). Systematic Entomology 31: 113–144.

[B57] Britton, E. B. 1957. A Revision of the Australian Chafers (Coleoptera: Scarabaeidae: Melolonthinae) vol. 1. British Museum (Natural History): London.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

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