Piratinae

Pirata hygrophilus, copyright Kjetil Fjellheim.

Belongs within: Lycosidae.

The Piratinae is a group of wolf spiders characterised by male pedipalps in which the functional conductor is combined with the short thin embolus into a sickle-shaped complex that rests in a deep, narrow ascending tegular groove (Yoo & Framenau 2006). Members of the genus Pirata, the pirate wolf spiders, are found in marshy habitats and may be seen running across water surfaces.

<==Piratinae
    |--Hygrolycosa Dahl 1908YF06
    |    `--H. rubrofasciata Ohlert 1865FY06
    |--Aulonia Koch 1847YF06
    |    |--*A. albimana (Walckenaer 1805)MGK03
    |    `--A. kratochvili Dunin, Buchar & Absolon 1986MGK03
    `--Pirata Sundevall 1833YF06
         |  i. s.: *P. piraticus (Clerck 1757)C01, MGK03 (see below for synonymy)
         |         P. insularis Emerton 1885MGK03
         |         P. latitans (Blackwall 1841)MGK03
         |         P. piratimorpha (Strand 1908) [=Tarentula piratimorpha, Piratula piratimorpha]J98
         |         P. piscatorius (Clerck 1757)K02 [=Lycosa (Pirata) piscatoriaK01, Tarentula (P.) piscatoriusS06]
         |         P. praedo (Kulcz. 1885) [=Tarentula (Pirata) praedo]S06
         |         P. raptor (Kulcz. 1885) [=Tarentula (Pirata) raptor]S06
         |         P. uliginosus (Thorell 1856)RKD02
         |--P. subpiraticus (Bösenberg & Strand 1906)PY07, YF06
         `--+--P. piratoidesPY07
            `--+--P. hygrophilus Thorell 1872PY07, VMP02
               `--P. knorri (Scopoli 1763)PY07, VMP02

*Pirata piraticus (Clerck 1757)C01, MGK03 [=Araneus piraticusS99, Lycosa (Pirata) piraticaK01, Tarentula (P.) piraticusS06]

*Type species of generic name indicated

References

[C01] Cambridge, F. O. P. 1901. A revision of the genera of the Araneae or spiders with reference to their type species. Annals and Magazine of Natural History, series 7, 7: 51–65.

[FY06] Framenau, V. W., & J.-S. Yoo. 2006. Systematics of the new Australian wolf spider genus Tuberculosa (Araneae: Lycosidae). Invertebrate Systematics 20: 185–202.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

[K02] Koponen, S. 2002. Ground-living spiders in bogs in northern Europe. Journal of Arachnology 30 (2): 262–267.

[K01] Kulczyński, V. 1901. Arachnoideák [Arachnoidea]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 311–369. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[MGK03] Marusik, Yu. M., E. F. Guseinov & S. Koponen. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) found in the country with description of three new species and brief review of Palaearctic Evippa Simon, 1885. Arthropoda Selecta 12 (1): 47–65.

[PY07] Park, Y. C., J.-S. Yoo, M. P. Schwarz, N. Murphy & J.-P. Kim. 2007. Molecular phylogeny of east Asian wolf spiders (Araneae: Lycosidae) inferred from mitochondrial 12S ribosomal DNA. Annals of the Entomological Society of America 100 (1): 1–8.

[RKD02] Relys, V., S. Koponen & D. Dapkus. 2002. Annual differences and species turnover in peat bog spider communities. Journal of Arachnology 30 (2): 416–424.

[S99] Simon, E. 1899. Ergebnisse einer Reise nach dem Pacific (Schauinsland 1896–1897). Arachnoideen. Zoologische Jahrbücher, Abteilung für Systematik, Geographie un Biologie der Thiere 12 (4): 411–437.

[S06] Strand, E. 1906. Die arktischen Araneae, Opiliones und Chernetes. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 431–478. Gustav Fischer: Jena.

[VMP02] Vink, C. J., A. D. Mitchell & A. M. Paterson. 2002. A preliminary molecular analysis of phylogenetic relationships of Australasian wolf spider genera (Araneae, Lycosidae). Journal of Arachnology 30 (2): 227–237.

[YF06] Yoo, J.-S., & V. W. Framenau. 2006. Systematics and biogeography of the sheet-web building wolf spider genus Venonia (Araneae: Lycosidae). Invertebrate Systematics 20: 675–712.

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