Emperor newt Tylototriton shanjing, copyright Spark.

Belongs within: Salamandridae.
Contains: Triturus.

The Pleurodelinae, newts, are a Holarctic group of salamanders with lives typically alternating between a terrestrial breeding and aquatic non-breeding phase.

| i. s.: Brachycormus [incl. Oligosemia]M00
| ProcynopsM00
| | `--Pleurodeles Michahelles 1830PW11, FG06
| | |--P. waltl Michaelles 1830PW11, W08
| | `--+--P. nebulosusPW11
| | `--P. poiretiPW11
| `--+--Chelotriton Pomel 1853M00, RB05
| | `--C. pliocenicus Bailon 1989B89
| |--Echinotriton Nussbaum & Brodie 1982PW11, FG06
| | |--E. andersoniPW11
| | `--E. chinhaiensisPW11
| `--Tylototriton Anderson 1871PW11, FG06
| | i. s.: T. andersoni (Boulenger 1892)I92
| | T. weigeltiM00
| |--T. asperrimusPW11
| `--+--T. wenxianensisPW11
| `--+--T. taliangensis Liu 1950PW11, L50
| `--+--T. shanjingPW11
| `--+--T. kweichowensis Fang & Chang 1932PW11, L50
| `--T. verrucosus Anderson 1871PW11, L50 [=T. verrocosaL50, T. verrucosaL50]
`--+--+--Notophthalmus Rafinesque 1820PW11, FG06
| | |--N. meridionalisPW11
| | `--+--N. perstriatusPW11
| | `--N. viridescens (Rafinesque 1820)PW11, W08 [=Triturus viridescensKSM06]
| `--Taricha Gray 1850PW11, FG06
| |--T. rivularisPW11
| `--+--T. granulosaPW11
| `--T. torosaPW11
`--+--Lissotriton Bell 1838PW11, FG06
| |--L. boscaiPW11
| `--+--+--L. helveticusPW11
| | `--L. italicusPW11
| `--+--L. montandoniPW11
| `--L. vulgaris (Linnaeus 1758)PW11, W08 [=Triturus vulgarisFG06]
| |--L. v. vulgarisK08
| `--‘Triturus’ v. meridionalisK08
`--+--Koaliella genzeli Herre 1950M00, M93
| | `--CalotritonPW11
| | |--C. arnoldiPW11
| | `--C. asper Dugès 1852PW11, FG06 [=Euproctus asperFG06]
| `--+--Ommatotriton Gray 1850PW11, FG06
| | |--O. ophryticus (Berthold 1846)PW11, W08 [=Triturus vittatus ophryticusW08]
| | `--O. vittatus (Jenyns 1835)PW11, RB05 [=Triturus vittatusFG06]
| `--Neurergus Cope 1862PW11, FG06
| |--N. strauchiiPW11
| `--+--N. kaiseriPW11
| `--+--N. crocatusPW11
| `--N. microspilotusPW11
`--+--Euproctus Gené 1838PW11, FG06 [incl. Megapterna Savi 1838FG06]
| |--E. montanus (Savi 1838)W08
| `--E. platycephalus (Gravenhorst 1829)W08
`--+--Ichthyosaura alpestrisPW11
|--Mesotriton Bolkay 1927FG06
| `--M. alpestris (Laurent 1768)FG06, W08 [=Triton alpestrisK08, Triturus alpestrisFG06]
`--+--Laotriton laoensisPW11
`--+--Pachytriton Boulenger 1878PW11, FG06
| |--P. brevipesPW11
| `--P. labiatusPW11
`--+--Cynops Tschudi 1838PW11, FG06
| |--+--C. ensicaudaPW11
| | `--C. pyrrhogaster (Boie 1826)PW11, W08
| `--+--C. cyanurusPW11
| `--+--C. orientalisPW11
| `--C. orphicusPW11
`--Paramesotriton Chang 1935PW11, FG06
|--+--P. caudopunctatusPW11
| `--P. zhijinensisPW11
`--+--P. chinensisPW11
`--+--P. hongkongensisPW11
`--+--P. deloustaliPW11
`--+--P. fuzhongensisPW11
`--P. guangxiensisPW11

*Type species of generic name indicated


[B89] Bailon, S. 1989. Les amphibiens et les reptiles du Pliocene superieur de Balaruc II (Herault, France). Palaeovertebrata 19 (1): 7–28.

[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[KSM06] King, R. C., W. D. Stansfield & P. K. Mulligan. 2006. A Dictionary of Genetics 7th ed. Oxford University Press.

[K08] Korsós, Z. 2008. History of the Herpetological Collection of the Hungarian Natural History Museum. Annales Historico-Naturales Musei Nationalis Hungarici 100: 37–93.

[L50] Liu, C.-C. 1950. Amphibians of western China. Fieldiana: Zoology Memoirs 2: 1–400.

[M93] Milner, A. R. 1993. Amphibian-grade Tetrapoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 665–679. Chapman & Hall: London.

[M00] Milner, A. R. 2000. Mesozoic and Tertiary Caudata and Albanerpetontidae. In: Heatwole, H., & R. L. Carroll (eds) Amphibian Biology vol. 4. Palaeontology. The evolutionary history of amphibians pp. 1412–1444. Surrey Beatty & Sons.

[PW11] Pyron, R. A., & J. J. Wiens. 2011. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61: 543–583.

[RB05] Rage, J.-C., & S. Bailon. 2005. Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France). Geodiversitas 27 (3): 413–441.

[W08] Warburg, M. R. 2008. Temporal variations in body dimensions followed for 25 years in a breeding population of adult salamanders, with a partial review on other Urodela (Amphibia, Urodela, Salamandridae). Senckenbergiana Biologica 88 (1): 81–101.

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