Polyptychitidae

Tollia tolli, from Ammoniten.org.

Belongs within: Dorsoplanitidae.
Contains: Polyptychitinae, Surites.

The Polyptychitidae are a morphologically diverse lineage of ammonites known from the Lower Cretaceous (Lower Ryazanian to Upper Hauterivian), with relatively simple sutures (Wright et al. 1996).

Polyptychitidae [Polyptichitidae]WCH96
    |--+--PolyptychitinaeWCH96
    |  `--+--Oosterellidae [Oosterellinae]WCH96
    |     |    |--Pseudosterella Spath 1924WCH96
    |     |    |    `--*P. fischeri (Nicklès 1892) [=Mortoniceras fischeri]WCH96
    |     |    `--Oosterella Kilian 1911 [=Nicklesia Kilian 1910 non Hyatt 1903]WCH96
    |     |         |--*O. cultrata (Orbigny 1841) [=Ammonites cultratus, *Nicklesia cultrata]WCH96
    |     |         `--O. steveniniWK81
    |     `--PlatylenticeratinaeWCH96
    |          |--Delphinites Sayn 1901 [incl. Proleopoldia Spath 1923, Pseudogarnieria Spath 1923]WCH96
    |          |    |--*D. ritteri Sayn 1901WCH96
    |          |    |--D. kurmyschensis (Stchirowsky 1894) [=Hoplites kurmyschensis, *Proleopoldia kurmyschensis]WCH96
    |          |    `--D. undulatoplicatile (Stchirowsky 1894) (see below for synonymy)WCH96
    |          |--Platylenticeras Hyatt 1900 [=Garnieria Sayn 1901 non Bourguignat 1877]WCH96
    |          |    |  i. s.: P. involutumWCH96
    |          |    |         P. robustumWCH96
    |          |    |--*P. (Platylenticeras) heteropleurum (Neumayr & Uhlig 1881) (see below for synonymy)WCH96
    |          |    `--P. (Tolypeceras Hyatt 1903)WCH96
    |          |         `--P. (*T.) marcousianum (Pictet & Campiche 1859) [=Ammonites marcousianus]WCH96
    |          `--Paquiericeras Sayn 1901WCH96
    |               |--*P. (Paquiericeras) paradoxum Sayn 1901WCH96
    |               |    |--P. p. paradoxumWCH96
    |               |    `--P. p. tuberculatumWCH96
    |               `--P. (Julianites Thieuloy 1977)WCH96
    |                    `--P. (*J.) undulatum Thieuloy 1977WCH96
    `--Tolliinae [Menjaitinae, Peregrinoceratidae, Suritidae, Suritinae, Tollinae]WCH96
         |--SuritesWCH96
         |--Thorsteinssonoceras Jeletzky 1965WCH96
         |    `--*T. ellesmerense Jeletzky 1965WCH96
         |--Peregrinoceras Sazonova 1971WCH96
         |    |--*P. pressulum (Bogolovskii 1897) [=Olcostephanus pressulus]WCH96
         |    `--P. albidumWCH96
         |--Menjaites Sazonova 1971 (see below for synonymy)WCH96
         |    |--*M. imperceptus Sazonova 1971WCH96
         |    |--M. certusWCH96
         |    |--*Costamenjaites’ jucundus Sazonova 1971WCH96
         |    |--*Sabydites’ menjaiteformis Bogomolov & Burdykina 1989WCH96
         |    `--*Luppoviceras’ validum Sazonova 1977WCH96
         |--Borealites Klimova 1969WCH96
         |    |--B. (Borealites) [incl. Pronjaites Sazonova 1971]WCH96
         |    |    |--*B. (B.) fedorovi Klimova 1969WCH96
         |    |    `--B. (B.) bidevexus (Bogoslovskii 1897) [=Olcostephanus bidevexus, *Pronjaites bidevexus]WCH96
         |    |--B. (Pseudocraspedites Casey, Mesezhnikov & Shul’Gina 1977)WCH96
         |    |    `--B. (*P.) anglicus (Shul’gina 1972) [=Subcraspedites anglicus]WCH96
         |    `--B. (Ronkinites Shul’gina 1972)WCH96
         |         `--B. (*R.) rossicus (Shul’gina 1972) [=Subcraspedites (*R.) rossicus]WCH96
         |--Virgatoptychites Voronets 1958WCH96
         |    |--V. (Virgatoptychites)WCH96
         |    |    |--*V. (V.) changalassensis Voronets 1958WCH96
         |    |    `--V. (V.) pachsaensisWCH96
         |    `--V. (Propolyptychites Kemper 1964) [incl. Paratollia Casey 1973]WCH96
         |         |--V. (*P.) quadrifidus (Koenen 1902) [=Polyptychites quadrifidus]WCH96
         |         |--*Paratollia’ kemperi Casey 1973WCH96
         |         `--V. (P.) pumilioWCH96
         |--Nikitinoceras Sokolov 1913 [=Temnoptychites Pavlow 1914]WCH96
         |    |  i. s.: ‘Temnoptychites’ syzranicusR81
         |    |--*N. (Nikitinoceras) hoplitoides (Nikitin 1888) [=Olcostephanus hoplitoides, *Temnoptychites hoplitoides]WCH96
         |    |--N. (Bodylevskiceras Sazonova 1977)WCH96
         |    |    `--N. (*B.) elegans (Bodylevskii in Luppov, Bodylevskii & Glazunova 1949) [=Temnoptychites elegans]WCH96
         |    `--N. (Russanovia Bodylevskii 1967) (see below for synonymy)WCH96
         |         |--N. (*R.) diptychum (Keyserling 1846) [=Ammonites diptychus]WCH96
         |         |--‘Temnoptychites (*Subtemnoptychites)’ prodigialis Klimova 1978WCH96
         |         `--N. (R.) rudis Bodylevskii 1967WCH96
         `--Tollia Pavlow 1914WCH96
              |  i. s.: T. complanatusWCH96
              |         T. undulatusWCH96
              |--T. (Tollia) [incl. Bodylevskites Klimova 1978, Neotollia Shul’gina 1969]WCH96
              |    |--*T. (T.) tolli Pavlow 1914WCH96
              |    |--*Bodylevskites’ harabylensis Klimova 1978WCH96
              |    `--T. (T.) klimovskiensis Krimgol’ts in Krimgol’ts, Petrova & Pchelintsev 1953 (see below for synonymy)WCH96
              `--T. (Neocraspedites Spath 1924)WCH96
                   `--T. (*N.) semilaevis (Koenen 1902) [=Craspedites semilaevis]WCH96

Delphinites undulatoplicatile (Stchirowsky 1894) [=Oxynoticeras undulatoplicatile, *Pseudogarnieria undulatoplicatile]WCH96

Menjaites Sazonova 1971 [incl. Costamenjaites Sazonova 1971, Luppoviceras Sazonova 1977, Sabydites Bogomolov & Burdykina 1989]WCH96

Nikitinoceras (Russanovia Bodylevskii 1967) [incl. Keyserlingyceras Sazonova 1961 (n. n.), Subtemnoptychites Klimova 1978]WCH96

*Platylenticeras (Platylenticeras) heteropleurum (Neumayr & Uhlig 1881) [=Amaltheus (Oxynoticeras) heteropleurus, *Garnieria heteropleura]WCH96

Tollia (Tollia) klimovskiensis Krimgol’ts in Krimgol’ts, Petrova & Pchelintsev 1953 [=*Neotollia klimovskiensis]WCH96

*Type species of generic name indicated

References

[R81] Rawson, P. F. 1981. Early Cretaceous ammonite biostratigraphy and biogeography. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 499–529. Academic Press.

[WK81] Wiedmann, J., & J. Kullmann. 1981. Ammonoid sutures in ontogeny and phylogeny. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 215–255. Academic Press.

[WCH96] Wright, C. W., J. H. Calloman & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

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