Potentilla

Creeping cinquefoil Potentilla reptans, copyright Olivier Pichard.

Belongs within: Roperculina.
Contains: Horkelia, Ivesia.

Potentilla, cinquefoils, is a northern temperate genus of herbs and shrubs with usually yellow flowers (Hickman 1993).

Characters (from Hickman 1993): Annual to shrub; odor resinous or absent. Leaves generally basal, odd-1-pinnate to 1-palmate or 1-ternate; leaflets more or less toothed or lobed, terminal generally more or less equal to lateral. Inflorescence a cyme, generally more or less open; pedicels generally more or less straight. Flower with hypanthium more or less shallow; bractlets 5; sepals generally 5, more or less triangular; petals generally 5, generally the same size as or larger than sepals, generally more or less widely obcordate, generally yellow; stamens generally 20; pistils generally many, styles generally jointed near tip. Fruit an achene.

<==Potentilla Linnaeus 1753 A61 [incl. Duchesnea EH03]
|–+–P. anserina EH03
|  |    |–P. a. ssp. anserina [incl. P. anserina var. sericea] H93
|  |    `–P. a. ssp. pacifica [incl. P. egedii var. grandis] H93
|  `–+–P. peduncularis EH03
|     `–P. stenophylla EH03
`–+–P. micrantha EH03
`–+–+–*P. reptans EH03
|  `–P. indica [=Duchesnea indica] EH03
`–+–P. dickinsii EH03
|–P. fragarioides EH03
`–+–+–P. chinensis EH03
|  |–P. multifida EH03
|  `–P. nivea EH03
`–+–P. norvegica EH03 [incl. P. norvegica ssp. monspeliensis H93]
`–+–Horkelia EH03
`–Ivesia EH03

Potentilla incertae sedis:
P. adenophylla H03
P. anglica H93
P. anserinoides Raoul 1844 [=P. anserina var. anserinoides (Raoul) Hooker 1852] A61
P. argentea BT72
P. argyrophylla O88
|–P. a. var. argyrophylla O88
`–P. a. var. atrosanguinea O88
P. biennis H93
P. biflora O88
P. borneensis H03
P. brassii H03
P. caliginosa O88
P. caulescens PS98
P. concinna [incl. P. beanii, P. concinna var. divisa] H93
P. coriandrifolia O88
P. cristae H93
P. cuneata O88
P. diversifolia [incl. P. glaucophylla] H93
P. drummondii H93
|–P. d. ssp. drummondii H93
|–P. d. ssp. breweri [incl. P. millefolia var. algida] H93
`–P. d. ssp. bruceae H93
P. erecta EH03
P. eriocarpa O88
P. exigua O88
P. flabellifolia H93
P. forrestii O88
|–P. f. var. forrestii O88
`–P. f. var. caespitosa O88
P. frigida C55a
P. fulgens O88
P. glandulosa D51
|–P. g. ssp. glandulosa [incl. P. glandulosa ssp. typica] H93
|–P. g. ssp. ashlandica [incl. P. glandulosa var. austiniae] H93
|–P. g. ssp. ewanii H93
|–P. g. ssp. globosa [incl. P. rhomboidea] H93
|–P. g. ssp. hansenii H93
|–P. g. ssp. nevadensis H93
|–P. g. ssp. pseudorupestris H93
`–P. g. ssp. reflexa H93
P. gracilis H93
|–P. g. var. gracilis H93
|–P. g. var. elmeri [incl. P. flabelliformis var. inyoensis, P. pectinisecta] H93
|–P. g. var. fastigiata (see below for synonymy) H93
|–P. g. var. flabelliformis H93
`–P. g. var. pulcherrima H93
P. grayi [=P. flabellifolia var. grayi] H93
P. heptaphylla S96
P. hickmanii H93
P. microphylla O88
|–P. m. var. microphylla O88
|–P. m. var. achilleifolia O88
`–P. m. var. commutata O88
P. millefolia [incl. P. millefolia var. klamathensis] H93
P. monanthes O88
P. morefieldii H93
P. multijuga H93
P. newberryi H93
P. pennsylvanica C55b [incl. P. pensylvanica var. strigosa H93]
P. polyphylla O88
P. pusilla S96
P. recta W69
P. rimicola [=P. wheeleri var. rimicola] H93
P. rivalis [incl. P. rivalis var. millegrana] H93
P. saundersiana O88
P. supina VB02
P. wheeleri H93

Potentilla gracilis var. fastigiata [incl. P. gracilis var. glabrata, P. gracilis ssp. nuttallii, P. gracilis var. permollis] H93

*Type species of generic name indicated

REFERENCES

[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[BT72] Baker, E. W., & D. M. Tuttle. 1972. New species and further notes on the Tetranychoidea mostly from the southwestern United States (Acarina: Tetranychidae and Tenuipalpidae). Smithsonian Contributions to Zoology 116: 1–37.

[C55a] Candolle, A. de. 1855a. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 1. Librairie de Victor Masson: Paris.

[C55b] Candolle, A. de. 1855b. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[D51] Dobzhansky, T. 1951. Genetics and the Origin of Species 3rd ed. Columbia University Press: New York.

[EH03] Eriksson, T., M. S. Hibbs, A. D. Yoder, C. F. Delwiche & M. J. Donoghue. 2003. The phylogeny of Rosoideae (Rosaceae) based on sequences of the internal transcribed spacers (ITS) of nuclear ribosomal DNA and the trnL/F region of chloroplast DNA. International Journal of Plant Science 164 (2): 197–211.

[H03] Heads, M. 2003. Ericaceae in Malesia: vicariance biogeography, terrane tectonics and ecology. Telopea 10 (1): 311–449.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: an introductory note. In: Ohba, H., & S. B. Malla (eds) The Himalayan Plants vol. 1. The University Museum, University of Tokyo, Bulletin 31: 19–46.

[PS98] Prosser, F., & S. Scortegagna. 1998. Primula recubariensis, a new species of Primula sect. Auricula Duby endemic to the SE Prealps, Italy. Willdenowia 28: 27–46.

[S96] Schatz, H. 1996. Oribatid mites (Oribatida) in alpine dry meadows (Austria, central Alps). In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 633–635. Ohio Biological Survey: Columbus (Ohio).

[VB02] Vijay, S. K., & T. N. Bhardwaja. 2002. Vegetation and phenodynamics of wetlands of central Rajasthan. Journal of the Bombay Natural History Society 99 (3): 573–581.

[W69] Wiens, J. A. 1969. An approach to the study of ecological relationships among grassland birds. Ornithological Monographs 8: 1–93.

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