Protoribates (Protoribates)

Protoribates lophothrichus, copyright R. Penttinen.

Belongs within: Haplozetidae.

The type subgenus of the genus Protoribates is a cosmopolitan group of oribatid mites (Subías 2004), albeit one of uncertain composition owing to confusion over the identity of the type species. True Protoribates have movable pteromorphs and had previously been referred to by the junior name Xylobates (Colloff & Halliday 1998).

Characters (from Balogh & Balogh 1992a, as Protoribates and Xylobates): Prodorsum with lamellae present, translamella absent; interlamellar setae long. Dorsosejugal suture present. Notogaster smooth; ten pairs of notogastral setae short but present; four pairs of areae porosae; areae porosae Aa, A2 and A3 round; pteromorphs well developed, movable. Four to six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs monodactyle.

<==Protoribates Berlese 1908 (Protoribates) (see below for synonymy)S04
    |--*P. (P.) dentatus (Berlese 1883)S04 (see below for synonymy)
    |--P. (P.) albidus (Ewing 1908)S04 [=Oribata albidaS04, Oribatula albidaCH98, *Propescheloribates albidusCH98]
    |--P. (P.) angulatus Choi 1986S04
    |--P. (P.) antillensis (Mahunka 1985) [=Xylobates antillensis]S04
    |--P. (P.) bayanicus Bayartogtokh 2000S04
    |--P. (P.) boraboraensis Sellnick 1959S04
    |--P. (P.) brevisetosus (Fujita 1989) [=Xylobates brevisetosus]S04
    |--P. (P.) capucinus Berlese 1908S04 [=Xylobates capucinusBB92]
    |    |--P. c. capucinusS04 (see below for synonymy)
    |    |--P. c. bonairensis Willmann 1936S04
    |    `--P. c. tentaculus Mihelčič 1958S04
    |--P. (P.) concretus (Kramer 1898) (n. d.) [=Oribata concreta]S04
    |--P. (P.) crassisetiger Choi 1986S04
    |    |--P. c. crassisetigerS04
    |    `--P. c. nipponicus Fujita 1989S04
    |--P. (P.) diani (Mahunka 1986) [=Xylobates diani]S04
    |--P. (P.) geonjiensis Choi 1994S04
    |--P. (P.) gracilis (Aoki 1982) [=Xylobates gracilis]S04
    |--P. (P.) hakonensis Aoki 1994S04
    |--P. (P.) halleri (Michael 1898) (n. d.) [=Oribata halleri, O. simplex Haller 1884 non Gervais 1849]S04
    |--P. (P.) imperfectus (Banks 1906) [=Galumna imperfecta]S04
    |--P. (P.) iracemae Pérez-Íñigo & Baggio 1994S04
    |--P. (P.) longiuscula (Koch 1841) (n. d.)S04 [=Zetes longiusculaS04, Notaspis longiusculaM98]
    |--P. (P.) lophothrichus (Berlese 1904)S04 (see below for synonymy)
    |--P. (P.) madagascarensis (Balogh 1961) [=Scheloribates madagascarensis]S04
    |--P. (P.) magnus Aoki 1982S04
    |--P. (P.) mollicoma (Hammer 1973) [=Xylobates mollicoma]S04
    |--P. (P.) myrmecophilus Aoki & Ito 1997S04
    |--P. (P.) nagaroensis (Fujita 1989) [=Xylobates nagaroensis]S04
    |--P. (P.) natalensis (Pletzen 1963) [=Scheloribates natalensis]S04
    |--P. (P.) ‘oblongus’ (Ewing 1909) (see below for synonymy)S04
    |--P. (P.) paracapucinus (Mahunka 1988) [=Xylobates paracapucinus]S04
    |--P. (P.) ‘parvulus’ Willmann 1931 non Berlese 1916S04
    |--P. (P.) pembertoni (Jacot 1934) [=Xylobates pembertoni]S04
    |--‘Xylobates’ punctulatus (Sellnick 1925)C-R79
    |--P. (P.) robustior (Jacot 1937) (see below for synonymy)S04
    |--P. (P.) rodriguezi (Mahunka 1988) [=Xylobates rodriguezi]S04
    |--P. (P.) rotundus (Aoki 2002) [=Xylobates rotundus]S04
    |--P. (P.) souchnaiensis Abdel-Hamid 1964S04 [=Xylobates souchnaiensisAFK91]
    |--P. (P.) tenuis (Wen, Aoki & Wang 1984) [=Xylobates tenuis]S04
    |--P. (P.) triangularis (Hammer 1971) [=Xylobates triangularis]S04
    |--P. (P.) varisetiger (Wen, Aoki & Wang 1984) [=Xylobates varisetiger]S04
    `--P. (P.) yezoensis (Fujikawa 1983) [=Xylobates yezoensis]S04

Protoribates Berlese 1908 (Protoribates) [incl. Alloribates Banks 1947, Propescheloribates Jacot 1936, Styloribates Jacot 1934, Xylobates Jacot 1929]S04

Protoribates (Protoribates) capucinus capucinus Berlese 1908S04 [incl. Xylobates capucinus angustior Jacot 1937S04, *Alloribates singularis Banks 1947CH98, S04]

*Protoribates (Protoribates) dentatus (Berlese 1883)S04 [=Oribata dentataS04, Oribates dentatusM98, Proribates (Scheloribates) dentatusB16; incl. Oribata monodactyla Haller 1884S04, Oribates monodactylusS59, Protoribates monodactylusB61, Xylobates monodactylusC-R79, Peloribates vastus Mihelčič 1956S04]

Protoribates (Protoribates) lophothrichus (Berlese 1904)S04 [=Oribata lophothrichaS04, Oribates lophotrichusCH98, *Xylobates lophotrichusCH98; incl. *Styloribates pectinatus Jacot 1934CH98, S04, Scheloribates pectinatusP63, Xylobates prionotus Woolley 1968S04]

Protoribates (Protoribates) ‘oblongus’ (Ewing 1909) [=Oribata oblonga non Koch 1879; incl. O. longa Ewing 1909, Xylobates longisetae Jacot 1937]S04

Protoribates (Protoribates) robustior (Jacot 1937) [=Xylobates oblongus robustior; incl. X. capucinus robustior Jacot 1937 non X. oblongus robustior Jacot 1937]S04

*Type species of generic name indicated


[AFK91] Alberti, G., N. A. Fernandez & G. Kümmel. 1991. Spermatophores and spermatozoa of oribatid mites (Acari: Oribatida). Part II: functional and systematical considerations. Acarologia 32 (4): 435–449.

[B61] Balogh, J. 1961. Identification keys of world oribatid (Acari) families and genera. Acta Zoologica Hungarica 1: 243–344.

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[C-R79] Corpuz-Raros, L. A. 1979. Philippine Oribatei (Acarina). I. Preliminary list of species and descriptions of forty new species. Philippine Agriculturist 62 (1): 1–82.

[M98] Michael, A. D. 1898. Oribatidae. In: H. Lohmann (ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen vol. 3. Acarina pp. 1–93. R. Friedländer und Sohn: Berlin.

[P63] Pletzen, R. van. 1963. Studies on South African Oribatei (Acarina). Acarologia 5 (4): 690–703.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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