Pselaphitae

 Centrophthalmus punctipennis, from here.

Belongs within: Pselaphinae.
Contains: Tmesiphorus, Pseudophanias, Stipesa, Arhytodini, Pselaphus, Tyraphus, Curculionellus, Clavigeritae, Ctenistes, Ctenisophus, Enoptostomus, Sognorus, Tyromorphus, Centrophthalmina, Tyrina, Hamotus.

The Pselaphitae are a large group of beetles that usually have eleven-segmented antennae and greatly developed maxillary palpi (Richards & Davies 1977).

<==Pselaphitae [Macroscelidae]
    |--+--+--ArhytodiniLC20
    |  |  `--+--Rhytus Westwood 1870LC20, R04b
    |  |     `--+--PselaphogeniusLC20
    |  |        `--PselaphiniR04a
    |  |             |--PselaphusNG05
    |  |             |--Dicentrius merkli Reitter 1879R04a
    |  |             |--Margaris imperialisR04a
    |  |             |--TyraphusR04a
    |  |             |--Neopselaphus Jeannel 1951NT01
    |  |             |    `--N. mexicanus (Park 1945)NT01
    |  |             |--Pselaphischnus Raffray 1897R04a
    |  |             |    `--P. squamosus Raffray 1897R04a
    |  |             |--Pselaphoxys Raffray 1890R04a
    |  |             |    `--P. delicatulus Raffray 1882R04a
    |  |             |--Mentraphus Sharp 1883R04a
    |  |             |    `--M. pselaphoides Sharp 1883R04a
    |  |             |--‘Psilocephalus’ Raffray 1877 non Swainson 1839R04a
    |  |             |    |--P. formicetorum Raffray 1877R04a
    |  |             |    `--P. lewisi Raffray 1887R04a
    |  |             |--Pselaphoptrus Reitter 1891R04a [=Pselaphopterus (l. c.)R05]
    |  |             |    |--P. banghaasi Reitter 1893R04a
    |  |             |    |--P. kubischteki Reitter 1891R04a
    |  |             |    `--P. lomnickii Reitter 1901R04a
    |  |             |--PselaphophusR04a
    |  |             |    |--P. atriventris (Westwood 1856) (see below for synonymy)R04a
    |  |             |    |--P. bicolorR04a
    |  |             |    |--P. sulciceps Raffray 1903R04a
    |  |             |    `--P. unicolor Raffray 1900R04a
    |  |             `--CurculionellusLC20
    |  `--ClavigeritaeLC20
    `--+--CtenistiniNG05
       |    |--CtenistesNG05
       |    |--CtenisophusR04a
       |    |--EnoptostomusR04a
       |    |--SognorusR04a
       |    |--Biotus Casey 1887R04a
       |    |    `--B. formicarius Casey 1887R04a
       |    |--Chenniopsis Raffray 1904R04a
       |    |    `--*C. madecassa Raffray 1904R04a
       |    |--Ctenisomimus Raffray 1904R04a
       |    |    `--*C. oneili (Raffray 1898) [=Sognorus oneili]R04a
       |    |--Narcodes King 1863 [incl. Edocranes]R05
       |    |    `--N. varia King 1863 [incl. N. pulchra King 1863]R05
       |    |--Atinus Horn 1868R04a
       |    |    |--A. brevicornis Casey 1893R04a
       |    |    `--A. monilicornis (Brendel 1866)R04a
       |    |--Ctenicellus Raffray 1904R04a
       |    |    |--C. laticollis Raffray 1900R04a
       |    |    `--C. major Raffray 1904R04a
       |    |--Laphidioderus Raffray 1887R04a
       |    |    |--L. brevipennis Raffray 1897R04a
       |    |    `--L. capensis Raffray 1887R04a
       |    |--Epicaris [incl. Taphrophorus Schaufuss 1882]R04a
       |    |    |--E. doriae Schaufuss 1882R04a
       |    |    `--E. ventralis Raffray 1882R04a
       |    |--Ctenisodes Raffray 1897LC20, NT01 [incl. Pilopius Casey 1897NT01]
       |    |    |--C. laticeps Raffray 1896R04a
       |    |    |--‘Pilopius’ ocularisC90
       |    |    `--C. piceusNT01
       |    |--Ctenisomorphus Raffray 1890R04a
       |    |    |--C. alternans Raffray 1892R04a
       |    |    |--C. elaniticus Raffray 1903R04a
       |    |    `--C. major Raffray 1877R04a
       |    |--Centrotoma Heyden 1849R04a
       |    |    |--C. bruckiR04a
       |    |    |--C. globulipalpusR04a
       |    |    |--C. lucifuga Heyden 1849R04a
       |    |    |--C. ludyiR04a
       |    |    |--C. penicillata Schaufuss 1863 [incl. C. rubra Saulcy 1864]R04a
       |    |    `--C. prodiga Sharp 1874R04a
       |    |--Ctenisis Raffray 1890R04a
       |    |    |--C. aequinoctialis Aubé 1844R04a
       |    |    |--C. amazonica Raffray 1896R04a
       |    |    |--C. angustata Raffray 1896R04a
       |    |    |--C. dispar (Sharp 1887) [=Desimia dispar; incl. C. brevicollis Raffray 1896]R04a
       |    |    |--C. nasuta Raffray 1896R04a
       |    |    `--C. raffrayi Casey 1893 [incl. C. dispar Raffray 1896 non Desimia dispar Sharp 1887]R04a
       |    |--Poroderus Sharp 1883R04a
       |    |    |--P. angusticeps (Raffray 1887) [=Enoptostomus angusticeps]R04a
       |    |    |--P. armatus Sharp 1874R04a
       |    |    |--P. biarmatus Raffray 1892R04a
       |    |    |--P. javanus [=Enoptostomus javanus]R04a
       |    |    |--P. medius Sharp 1874R04a
       |    |    |--P. siamensis [=Enoptostomus siamensis]R04a
       |    |    `--P. similis Sharp 1874R04a
       |    |--Chennium Latreille 1807R04a
       |    |    |--C. antennatum Reitter 1881R04a
       |    |    |--C. bituberculatum Latreille 1807R04a
       |    |    |--C. eppelsheimi Reitter 1887R04a
       |    |    |--C. judaeumR04a
       |    |    |--C. kiesenwetteriR04a
       |    |    |--C. paulinoi Reitter 1887R04a
       |    |    |--C. prometheus Saulcy 1875R04a
       |    |    |--C. semenowi Reitter 1891R05
       |    |    `--C. steigerwaldi Reitter 1881R04a
       |    `--DesimiaR04a
       |         |--D. arabica Raffray 1887R04a
       |         |--D. caviceps Raffray 1904R04a
       |         |--D. darius [=Tmesiphorus darius]R04a
       |         |--D. frontalis Raffray 1887 [incl. D. depilis Raffray 1887]R04a
       |         |--D. ghilianii Aubé 1844 [=Tmesiphorus ghilianii; incl. D. complex, D. integricollis Fairmaire 1858]R04a
       |         |--D. gibbicollis Raffray 1899R04a
       |         |--D. parvipalpis Raffray 1873R04a [incl. D. pici Croiss. 1894R05]
       |         |--D. sharpi Raffray 1883R04a
       |         `--D. subcalva Reitter 1882R04a
       `--TyriniNG05
            |  i. s.: ‘Batrisus’ angulatus Westw. 1856R04a
            |         TyropsisR04a
            |           |--T. adumbrata Reitter 1885R04b
            |           |--T. castanea (see below for synonymy)R04b
            |           |--T. cavifrons Raffray 1895 [incl. T. valdiviensis Schaufuss 1886 non Reitter 1885]R04b
            |           |--T. difformisR04b
            |           |--T. hirta Reitter 1888R04b
            |           |--T. palpalis Reitter 1883 [incl. T. praeses]R04b
            |           |--T. pilifera Reitter 1888R04b
            |           |--T. spinula Reitter 1885R04b
            |           `--T. valdiviensis Reitter 1885 non Pselaphus valdiviensis (not preoc. if in dif. gen.)R04b
            |         Somatipion globuliferR04b
            |         Centrophthalmosis Raffray 1904R04a
            |           |--C. barbatus (Reitter 1882) [=Centrophthalmus barbatus]R04b
            |           |--C. exilis (Raffray 1882) (see below for synonymy)R04b
            |           |--C. inexpectus Raffray 1904R04a
            |           |--C. monilis Raffray 1882R04b
            |           |--C. muticus Raffray 1898R04b
            |           |--C. reitteri Raffray 1904R04a
            |           `--C. sulcatus Raffray 1904R04a
            |         Acylopselaphus Raffray 1883R04a, R04b
            |           |--A. alluaudi Raffray 1903R04b
            |           |--A. calcaratus Raffray 1897R04b
            |           |--A. clavatus Raffray 1903R04b
            |           |--A. communis (Schaufuss 1890) [=Centrophthalmus communis]R04b
            |           |--A. crassicornis Raffray 1903R04b
            |           |--A. madagascariensis Raffray 1903R04b
            |           `--A. mariae Raffray 1883R04b
            |         Zeatyrus Sharp 1881R04a, R04b [incl. Parmipalpus Broun 1881R04b]
            |           `--Z. lawsoni Sharp 1881 [incl. Z. montivagus Broun 1881]R04b
            |         Leanymus Raffray 1900R04a, R04b
            |           |--L. mirus Lea 1919L19
            |           `--L. palpalis Raffray 1900R04b
            |         Pselaphocerus Raffray 1890R04a, R04b
            |           |--P. acutispina Raffray 1898R04b
            |           |--P. amicus Raffray 1898R04b
            |           |--P. antennatus Raffray 1897R04b
            |           |--P. diversus Raffray 1897R04b
            |           |--P. heterocerus Raffray 1887R04b
            |           |--P. nodicornis Raffray 1898R04b
            |           `--P. peringueyi Raffray 1887R04b
            |         SintectodesR04a
            |           |--S. crassus Schaufuss 1887R04b
            |           |--S. diversipalpus Reitter 1885R04b
            |           `--S. tortipalpus Raffray 1901R04b
            |         Eulasinus Sharp 1892R04a, R04b
            |           `--E. walkeri Sharp 1892R04b
            |         Subulipalpus Schaufuss 1877R04a, R04b
            |           |--S. myrmecophilus Raffray 1904R04a
            |           `--S. spinicoxis Schaufuss 1877R04b
            |         Marellus Motschulsky 1851R04a, R04b
            |           |--M. aegyptiacus Motsch. 1851R04b
            |           |--M. biskrensis Reitter 1884R04b
            |           |--M. filipalpis Raffray 1887R04b
            |           |--M. granosus Raffray 1897R04b
            |           `--M. palpator Raffray 1877R04b
            |         Didymoprora Raffray 1890R04a, R04b
            |           |--D. armata Raffray 1900R04b
            |           |--D. dimidiata Raffray 1900R04b
            |           |--D. leana Raffray 1900R04b
            |           |--D. puncticollis Raffray 1900R04b
            |           |--D. semipunctata Raffray 1904R04a
            |           `--D. victoriae King 1865R04b [=Tyrus victoriaeM96]
            |         Palimbolus Raffray 1900R04a, R04b
            |           `--*P. mirandus (Sharp 1874)R04a, R04b [=Tyrus mirandusR04a]
            |         Spilorhombus Raffray 1900R04a, R04b
            |           `--S. hirtus Raffray 1900R04b
            |         Lethenomus Raffray 1895R04a, R04b
            |           `--L. villosus [=Aplodea villosa]R04b
            |         Tyrogetus Broun 1893R04a, R04b
            |           `--T. optandus Broun 1893R04b
            |         Neotyrus Raffray 1895R04a, R04b
            |           `--N. gibbicollis [=Aplodea gibbicollis]R04b
            |         Schaufussia Raffray 1883R04a, R04b
            |           |--S. formosa (King 1863) (see below for synonymy)R04b
            |           `--S. nasuta Raffray 1904R04a
            |         Gerallus Sharp 1874 [incl. Zozimus]R04b
            |           |--G. cribratipennisR04b
            |           |--G. dimidiatus Raffray 1900R04b
            |           |--G. longipesR04b
            |           |--G. modestusR04b
            |           |--G. nanus Sharp 1874R04b
            |           |--G. palpalis King 1863R04b
            |           `--G. punctipennis [incl. G. globulicornis]R04b
            |         Hamotulus [incl. Tychotyrus Broun 1893]R04b
            |           |--H. armatus Broun 1893R04b
            |           |--H. chamaeleonR04b
            |           |--H. curvipes Broun 1893R04b
            |           |--H. dispar Raffray 1900R04b
            |           |--H. mutandus (Sharp 1874) [=Tyrus mutandus]R04b
            |           |--H. spinipes Broun 1893R04b
            |           `--H. sternalis Broun 1893R04b
            |         Abascantus sannio Schauf. 1886R04b, M96
            |         TyromorphusR04b
            |         Taphrostethus Schaufuss 1882R04b
            |           |--T. clavatus (Raffray 1882) [=Tyrus clavatus]R04b
            |           |--T. hamotoides Schaufuss 1882R04b
            |           |--T. javanicus (Raffray 1882) [=Tyrus javanicus]R04b
            |           `--T. longispina Raffray 1903R04b
            |         Aploderina Raffray 1904R04a
            |           `--*A. sulcicornis Raffray 1904R04b
            |         Hamotopsis Raffray 1900R04b [=Homatopsis (l. c.)R05]
            |           `--H. australasiae Raffray 1900R04b
            |         Apharus Reitter 1882R04b
            |           |--A. armipes Raffray 1891R04b
            |           `--A. muelleri Reitter 1882R04b
            |         Cercoceropsis Raffray 1904R04a
            |           `--*C. longipes Raffray 1904R04b
            |         Pseudohamotus Raffray 1890R04b
            |           |--P. conjunctus Reitter 1882R04b
            |           |--P. curtipalpis Sharp 1887 [=Hamotus curtipalpus]R04b
            |           |--P. inflatipalpus Reitter 1888R04b
            |           |--P. latipalpis Sharp 1887 [=Hamotus latipalpus]R04b
            |           `--P. planiceps Raffray 1904R04b
            |         Cercoceroides Raffray 1896R04b
            |           |--C. germaini Raffray 1890R04b
            |           |--C. simplex Raffray 1904R04b
            |           `--C. tuberculatus Raffray 1904R04b
            |         Cercocerulus Raffray 1904R04a
            |           `--*C. hirsutus [=Phalepsus hirsutus]R04b
            |         Pseudotychus Raffray 1897R04b
            |           `--P. nigerrimus Raffray 1897R04b
            |         Eudranes Sharp 1892R04b
            |           `--E. carinatus Sharp 1892R04b
            |         Lasinus Sharp 1874LC20, R04b
            |           |--L. mandarinus Raffray 1890R04b
            |           |--L. mikadoLC20
            |           `--L. spinosus Sharp 1874R04b
            |         Durbos Sharp 1874R04b
            |           |--D. cribratipennis Schauf. 1886M96
            |           |--D. interruptus Schauf. 1886M96
            |           `--D. priscus Sharp 1874R04b
            |         Horniella Raffray 1904 [=Hornia Raffray 1901 (preoc.)]R04b
            |           `--H. hirtella Raffray 1901R04b
            |         Phamisulus Reitter 1888 [incl. Neophamisus Reitter 1888]R04b
            |           `--P. horroris [=Phamisus horroris, Neophamisus horroris; incl. Phamisulus hetschkoi Reitter 1888]R04b
            |         Rytus King 1866R04b
            |           |--R. corniger King 1865 [incl. R. porcellus]R04b
            |           |--R. emarginatus King 1866 [incl. R. victoriae King 1866]R04b
            |           |--R. gemmifer Schaufuss 1886R04b
            |           |--R. howarthi Raffray 1903R04b
            |           |--R. procurator Schauf. 1886M96 [incl. R. orientalis Schauf. 1886R04b, M96]
            |           |--R. setiger Raffray 1903R04b
            |           `--R. subulatus King 1865 [incl. R. punctatus King 1866, R. subasper]R04b
            |         Anagonus Fauvel 1903R05, R04a
            |           `--A. fracticornis Fauvel 1903R04a
            |--CentrophthalminaNG05
            |--TyrinaNG05
            `--HamotinaNT01
                 |--HamotusNT01
                 `--Circocerus Motschulsky 1855 [=Cercocerus (l. c.); incl. Upoluna Schaufuss 1886]NT01
                      `--C. batrisioides Motschulsky 1856NT01 [incl. Upoluna flavitarsis Schaufuss 1886R04b]
Pselaphitae incertae sedis:
  Schistodactylus Raffray 1883R04b [SchistodactyliniR03]
    `--S. phantasma Raffray 1883R04b
  Ceophyllus LeConte 1849NT01 [=Ceophillus (l. c.)R05; CeophylliniC90]
    `--C. monilis LeConte 1849NT01
  Phalepsus Westwood 1870R04b [=Phalespus (l. c.)R04b; PhalepsiniNG05]
    |--P. ampliventrisR04b
    |--P. batesellus Westwood 1870R04b
    |--P. cavicornis Raffray 1904R04b
    |--P. fluminicolaR04b
    |--P. marelloides Reitter 1888R04b
    |--P. nanusR04b
    |--P. subglobosus Westwood 1870R04b
    `--P. vulgaris Raffray 1904R04b
  Odontalgus Raffray 1877R04a [OdontalginiNG05]
    |--O. costatus Raffray 1897R04a
    |--O. gracilis Raffray 1904R04a
    |--O. longicornis Raffray 1898R04a
    |--O. palustris Raffray 1887R04a
    |--O. raffrayi Reitter 1882R04a
    |--O. tuberculatus Raffray 1877R04a
    |--O. validus Raffray 1904R04a
    |--O. vespertinus Raffray 1877R04a
    `--O. vestitus Schaufuss 1886NG05
  TmesiphoriniNG05
    |--TmesiphorusNG05
    |--PseudophaniasNG05
    |--Ctenotillus Raffray 1896NG05, R04b
    |    `--C. costatus Raffray 1896R04b
    |--Chandleriella Hlavac 2000NG05
    |    `--*C. termitophila (Bryant 1915) [=Lasinus termitophilus]NG05
    |--Raphitreus Sharp 1883NG05, R04b
    |    |--R. dentimanus Raffray 1890NG05
    |    `--R. speartus Sharp 1874R04b
    |--DacnotillusNG05
    |    |--D. pilosus Raffray 1895NG05
    |    |--D. transversus Raffray 1895NG05
    |    `--D. tuberculatus Raffray 1895NG05
    `--Ancystrocerus Raffray 1893NG05, R04b
         |--A. carinatus Raffray 1904R04a
         |--A. laevipennis Raffray 1896R04b
         |--A. longicornis Raffray 1895NG05
         |--A. militaris Raffray 1896R04b
         |--A. pallidus Raffray 1895NG05
         |--A. punctatus Raffray 1895NG05
         |--A. rugicollis Raffray 1895NG05
         `--A. sumatrensis Raffray 1893R04b
  HybocephaliniNG05
    |--StipesaNG05
    |--Pseudapharina Raffray 1890R04a
    |    `--P. prima (Reitter 1883)NG05 [=Filiger primaR04a]
    |--Metaxoides Schaufuss 1877R04a
    |    `--M. bruchiformis Schaufuss 1877R04a
    |--Apharinodes Raffry 1890R04a
    |    |--A. miranda Raffray 1895NG05
    |    `--A. squamosa Raffray 1890NG05
    |--Mestogaster Schmidt 1838R04a
    |    |--M. crassicornis Schmidt 1838R04a
    |    `--M. nitidicollis Reitter 1882R04a
    |--HybocephalodesNG05
    |    |--H. dentiventris Reitter 1883NG05
    |    |--H. informis Reitter 1883NG05
    |    `--H. telangensis Reitter 1883NG05
    |--Ephimia Reitter 1883R04a
    |    |--E. crassicornis Sharp 1887R04a
    |    |--E. simoni Reitter 1883R04a
    |    `--E. subnitida Raffray 1904R04a
    |--Hybocephalus Motsch. 1851R04a
    |    |--H. dentiventris Reitter 1883R04a
    |    |--H. informis Reitter 1883R04a
    |    |--H. minimus Motsch. 1851R04a
    |    |--H. squamosus Motsch. 1851R04a
    |    `--H. telangensis Reitter 1883R04a
    `--Apharina Reitter 1882R04a
         |--A. armata Raffray 1904R04a
         |--A. borneensis Raffray 1896NG05
         |--A. conicicollis (Schaufuss 1877) [=Filiger conicicollis]R04a
         |--A. fuscipennis Reitter 1883NG05
         |--A. nodula Raffray 1904R04a
         |--A. simoni Reitter 1882R04a
         `--A. squamiceps [=Panaphantus squamiceps]R04a
Nomina nuda: Cercocerus perplexus Motschulsky 1855R04b
             Desimia abyssinica Raffray 1904R04a
             Taphrostethus sumatrensis Raffray 1904R04b

Centrophthalmosis exilis (Raffray 1882) [=Centrophthalmus exilis; incl. Centrophthalmus grandipalpis Raffray 1882]R04b

Pselaphophus atriventris (Westwood 1856) [=Bryaxis atriventris; incl. P. anopunctatus, P. clavatus King 1865, P. clavicornis, P. atriventris var. edwardsi King 1865, P. atriventris var. simplicior, P. atriventris var. vexator]R04a

Schaufussia formosa (King 1863) [=Tyrus formosus; incl. Durbos affinis Schaufuss 1886, Schaufussia angustior Raffray 1883, Bryaxis brevis Schaufuss 1880, Tyromorphus constrictinasus Schaufuss 1886, Durbos intermedia Schaufuss 1886]R04b

Tyropsis castanea [=Pselaphus castaneus; incl. T. chevrolati, T. cosmoptera Schaufuss 1886 non Pselaphus cosmopterus (not preoc. if in dif. gen.), T. elsbethae Reitter 1883]R04b

*Type species of generic name indicated

References

[C90] Chandler, D. S. 1990. Insecta: Coleoptera Pselaphidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1175–1190. John Wiley & Sones: New York.

[L19] Lea, A. M. 1919. Notes on some miscellaneous Coleoptera with descriptions of new species.—Part V. Transactions of the Royal Society of South Australia 43: 166–261, pls 25–27.

[LC20] Lü, L., C.-Y. Cai, X. Zhang, A. F. Newton, M. K. Thayer & H.-Z. Zhou. 2020. Linking evolutionary mode to palaeoclimate change reveals rapid radiations of staphylinoid beetles in low-energy conditions. Current Zoology 66 (4): 435–444.

[M96] Masters, G. 1896. Catalogue of the described Coleoptera of Australia. Supplement, part II. Dytiscidae, Gyrinidae, Hydrophyllidae, Staphylinidae, Pselaphidae, Paussidae, Silphidae, Scaphididae, Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydiidae, Cucujidae, Cryptophagidae, Lathridiidae, Mycetophagidae, Dermestidae, Byrrhidae, Parnidae, Heteroceridae. Proceedings of the Linnean Society of New South Wales 21 (Suppl.): 695–754.

[NT01] Newton, A. F., M. K. Thayer, J. S. Ashe & D. S. Chandler. 2001. Staphylinidae Latreille, 1802. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 272–418. CRC Press: Boca Raton.

[NG05] Nomura, S., & I. Abd Ghani. 2005. Faunistic notes on the pselaphine species of the supertribes Goniaceritae, Pselaphitae and Clavigeritae from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga 10 (1–2): 1–36.

[R03] Raffray, A. 1903. Genera et catalogue des psélaphides. Annales de la Société Entomologique de France 72: 484–604.

[R04a] Raffray, A. 1904a. Genera et catalogue des psélaphides [part II]. Annales de la Société Entomologique de France 73: 1–400.

[R04b] Raffray, A. 1904b. Genera et catalogue des psélaphides [part III]. Annales de la Société Entomologique de France 73: 401–476, pls 1–3.

[R05] Raffray, A. 1905. Genera et catalogue des psélaphides [part IV]. Annales de la Société Entomologique de France 73: 635–658.

Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

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