Pseudocaeciliinae

Heterocaecilius lachlani, from Insects of Tasmania.

Belongs within: Pseudocaecilioidea.

The Pseudocaeciliinae are a group of barklice with two-segmented tarsi, known predominantly from south-east Asia and the Pacific (New & Lienhard 2007).

<==Pseudocaeciliinae [Meplerinae, Pseudoscottiinae] NL07
|–*Allopsocus marginalis Banks 1920 NL07
|–Scottiella NL07
|–Platyocaecilius Li 2002 L02b
|    `–*P. parallelivenius Li 2002 L02b
|–Fashengocaecilius Lienhard 2003 L03 [=Calocaecilius Li 2002 non Mockford 1974 L02b]
|    `–*F. biaduncus (Li 1992) [=Allocaecilius biaduncus, *Calocaecilius biaduncus] L03
|–Phyllocaecilius Li 2002 L02b
|    `–*P. atrichus Li 2002 L02b
|–Chorocaecilius Li 2000 L02a
|    |–C. brunellus (Tillyard 1923) (see below for synonymy) SN08
|    `–C. decemdentatus Li 2000 L02a
|–Phallocaecilius Lee & Thornton 1967 L02b
|    |–*P. hirsutus (Thornton 1961) [=Pseudocaecilius hirsutus] NL07
|    `–P. sentosus Li 1999 L02a
|–Thelocaecilius Li 2002 L02b
|    |–*T. mecokeratus Li 2002 L02b
|    `–T. papillatus (Li 1995) [=Heterocaecilius papillatus] L02b
|–Neocaecilius Li 2000 L02a
|    |–*N. campanula [=Heterocaecilius campanula] NL07
|    |–N. granulatus (Lee & New 1992) NL07
|    |–N. mutatus (Lee & New 1992) NL07
|    `–N. qianshanensis Li 2000 L02a
|–Mesocaecilius Okamoto 1910 NL07, L02a [Mesocaeciliinae]
|    |–*M. quadrimaculatus Okamoto 1910 L02a
|    |–M. bellus Li 2002 L02b
|    |–M. decorosus Li 2002 L02b
|    |–M. elegans Li 1997 L02b
|    |–M. euryopterus Li 2002 L02b
|    |–M. lepidus Li 2002 L02b
|    `–M. paraornatus (New 1977) [=Pseudocaecilius paraornatus] L02b
|–Ophiodopelma Enderlein 1908 L02a
|    |–*O. ornatipenne Enderlein 1908 NL07, L02a
|    |–O. anocella Li 2002 L02b
|    |–O. fluctosa Li 1995 L02a
|    |–O. glyptocephalum Lienhard 1985 NL07
|    |–O. hieroglyphicum (Enderlein 1903) NL07
|    |–O. multipunctatum (Hagen 1859) NL07
|    |–O. permaculatum Lee & Thornton 1967 NL07
|    |–O. pictipenna Lee & Thornton 1967 NL07
|    |–O. polyspila Li 1993 L02b
|    `–O. semiceps Lee & Thornton 1967 NL07
|–Levucaecilius Smithers 1995 NL07, S95
|    |–*L. greenwoodi [=Pseudocaecilius greenwoodi, Orbocaecilius greenwoodi] NL07
|    |–L. albicrus (Thornton 1981) [=Heterocaecilius albicrus] S95
|    |–L. apicalis (Thornton 1981) [=Heterocaecilius apicalis] S95
|    |–L. dardanus (Lee & Thornton 1967) S95 [=Heterocaecilius dardanus NL07, Neocaecilius dardanus NL07]
|    |–L. hilli Smithers 1995 NL07
|    |–L. panicus (Lee & Thornton 1967) [=Heterocaecilius panicus] S95
|    |–L. pictus (Thornton 1981) [=Heterocaecilius pictus] S95
|    |–L. simplex (Lee & Thornton 1967) NL07, L02a [=Heterocaecilius simplex NL07]
|    |–L. veitchi [=Heterocaecilius veitchi] S95
|    `–L. volatus (Lee & Thornton 1967) [=Heterocaecilius volatus] S95
|–Lobocaecilius Lee & Thornton 1967 L02a
|    |–*L. cynara Lee & Thornton 1967 L02a
|    |–L. arcuatus Vaughan, Thornton & New 1989 NL07
|    |–L. bifurcus Li 1993 L02a
|    |–L. drepanus New & Thornton 1976 NL07
|    |–L. ligur Lee & New 1992 NL07
|    |–L. longifurca Lee & Thornton 1967 NL07
|    |–L. pahangensis Lee & New 1992 NL07
|    |–L. perductivirgus Li 2002 L02b
|    |–L. pilosus Lee & New 1992 NL07
|    |–L. quadripartitus Li 1995 L02b
|    |–L. thorntoni Lee & New 1992 NL07
|    `–L. viridis Lee & New 1992 NL07
|–Allocaecilius Lee & Thornton 1967 L02a
|    |–*A. heterothrax Lee & Thornton 1967 L02a
|    |–A. albidorsualis Li 1993 L02a
|    |–A. arcuatus Li 2002 L02b
|    |–A. elongatus Lee & Thornton 1967 NL07
|    |–A. fusciceps Lee & Thornton 1967 NL07
|    |–A. gombakensis Lee & New 1992 NL07
|    |–A. guangdongicus Li 1993 L02a
|    |–A. maculatus Lee & Thornton 1967 NL07
|    |–A. selangorensis New & Thornton 1975 NL07
|    |–A. sinensis Lee & Thornton 1967 L02a
|    |–A. smithersi Lee & New 1992 NL07
|    `–A. tenuilongus Li 1995 L02a
|–Mepleres Enderlein 1926 SN08 [incl. Meniscopsocus Li 1993 L02b, Pseudoscottiella Badonnel 1946 L02b]
|    |–*M. maeandricus Enderlein 1926 SN08
|    |–M. cornutus (Lee & New 1992) NL07 [=Pseudoscottiella cornuta L02b]
|    |–‘Pseudoscottiella’ decolor Badonnel 1955 B55
|    |–‘Pseudoscottiella’ hyalina Badonnel 1955 B55
|    |–‘Pseudoscottiella’ immaculata Badonnel 1955 B55
|    |–M. indicus (Lee & New 1992) NL07
|    |–M. limbatus (Enderlein 1908) L02a
|    |–M. longicellus Li 2002 L02b
|    |–M. longitudinalis Li 2002 [=M. longitudianlis] L03
|    |–‘Pseudoscottiella’ maculata Broadhead & Richard 1982 L02b
|    |–*Pseudoscottiella’ megops NL07
|    |–M. nigroguttatus (Karny 1925) NL07 [=Pseudoscottiella nigraguttata L02b]
|    |–M. ocellatus Li 2002 L02b
|    |–M. oresbius Li 2002 L02b
|    |–M. parvicellus Li 2002 L02b
|    |–M. pisiformis (Lee & New 1992) NL07
|    |–M. plenimacularis Li 2002 L02b [=Pseudoscottiella plenimacularis L02a]
|    |–M. proboscideus Li 2002 L02b
|    |–M. procurrens Li 2002 L02b
|    |–M. radialis (Thornton 1984) NL07
|    |–M. raphalis (Lee & New 1992) NL07
|    |–M. rotundatus (New 1974) [=Pseudoscottiella rotundata, Meniscopsocus rotundatus] SN08
|    |–M. similis (Lee & New 1992) NL07
|    |–M. sinicus (Li 1993) L02b [=*Meniscopsocus sinicus NL07]
|    |–M. solenocerus (Lee & New 1992) NL07
|    |–M. suella (Thornton 1984) NL07 [=Pseudoscottiella suella L02b]
|    |–M. tanei (Smithers 1977) [=Pseudoscottiella tanei, Meniscopsocus tanei] SN08
|    |–M. transversus (Banks 1937) NL07
|    |–‘Pseudoscottiella’ tuberculata Badonnel 1955 B55
|    |–M. ulterior Li 2002 L02b
|    |–M. unitus Li 2002 L02b
|    |–M. urbanus (Vaughan, Thornton & New 1991) NL07
|    |–M. vilus (Lee & New 1992) NL07
|    |–‘Pseudoscottiella’ yeni New 1974 L02b
|    `–M. yunnanicus Li 2002 L02b
|–Pseudocaecilius Enderlein 1903 Y02 [incl. Hageniella Enderlein 1903 (preoc.) L02a]
|    |–P. citricola (Ashmead 1879) NL07 (see below for synonymy)
|    |–P. angustus (Enderlein 1903) NL07
|    |–P. bibulbus Li 1993 L03, L02a [=P. bibalbus (l. c.) L03]
|    |–P. bicostatus Li 2002 L02b
|    |–P. ceratocercus Li 2002 L02b
|    |–P. claggi (Banks 1937) NL07
|    |–P. clunialis Vaughan, Thornton & Neo 1991 NL07
|    |–P. cornutus Lee & Thornton 1967 NL07
|    |–P. danaus Lee & New 1992 NL07
|    |–P. euryocercus Li 2002 [=P. eurocerus] L03
|    |–P. formosanus (Banks 1937) L02a
|    |–P. galactozonalis Li 2002 [=P. galactozonacis] L03
|    |–P. helicoides Lee & Thornton 1967 NL07
|    |–P. immaculatus Li 2002 L02b
|    |–P. inaequalis (Banks 1916) NL07
|    |–P. innotatus Banks 1937 NL07
|    |–P. kagoshimensis Y02
|    |–P. largicellus Li 2002 L02b
|    |–P. machadoi Badonnel 1955 B55
|    |–P. monotaeniatus Li 2002 [=P. monotaenitus, P. monotaenius] L03
|    |–P. nitoris (Banks 1937) NL07
|    |–P. octomaculatus (Li 1995) [=Heterocaecilius octomaculatus] L02b
|    |–P. otiosus (Banks 1937) NL07
|    |–P. papillaris Li 2002 L02b
|    |–P. plagiozonalis Li 1995 L02a
|    |–P. productus New & Thornton 1976 NL07
|    |–P. pusillus (Banks 1931) NL07
|    |–P. ranus Thornton 1984 NL07
|    |–P. serratus Lee & New 1992 NL07
|    |–P. setifer Vaughan, Thornton & New 1991 NL07
|    |–P. sexdentatus Li 2002 L02b
|    |–P. tahitiensis (Karny 1926) [=Epipsocus tahitiensis] M93
|    |–P. tenellus Enderlein 1926 NL07
|    |–P. testaceus Enderlein 1903 NL07
|    |–P. undecimimaculatus Li 1995 L02b
|    |–P. utricularis Li 1999 L02a
|    |–P. venimaculatus Li 2002 L02b
|    |–P. wellsae Lee & New 1992 NL07
|    `–‘Psocus’ zonatus [=*Hageniella zonata] NL07
`–Heterocaecilius Lee & Thornton 1967 L02a [incl. Orbocaecilius Li 1999 L03, Obrocaecilius (l. c.) L03]
|–*H. minutus Lee & Thornton 1967 SN08 [=H. minotus NL07]
|–H. abacutidens Li 2002 L02b
|–H. analis Vaughan, Thornton & New 1991 NL07
|–H. anareolatus New 1993 NL07
|–H. anomalus (Thornton 1961) Y02, L02b [=Orbocaecilius anomolus L02b]
|–H. argutus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) argutus] L03
|–H. bicruris (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bicruris] L03
|–H. bidigitatus Li 1992 L02a [=Orbocaecilius bidigitatus L03]
|–H. bifarius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bifarius] L03
|–H. brachystigmus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) brachystigmus] L03
|–H. brevifurca Thornton 1984 NL07
|–H. challengerius Lee & New 1992 NL07
|–H. circulicellus Li 1995 L02b [=Orbocaecilius circulicellus L03]
|–H. dasoceratus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) dasoceratus] L03
|–H. deloneurous Li 2002 L02b
|–H. dentalis Li 2002 L02b
|–H. duodecidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) duodecidentus] L03
|–H. fasciatus Lee & Thornton 1967 NL07
|–H. flaviformis Lee & New 1992 NL07
|–H. fumosus Thornton 1984 NL07
|–H. fuscipalpus Lee & Thornton 1967 NL07
|–H. fuscus Y02
|–H. gressitti Lee & Thornton 1967 NL07
|–H. hepaticivenus Li 2002 L02b
|–H. hoplites Lee & New 1992 NL07
|–H. huananensis Li 2002 L02b
|–H. huaxiensis Li 2002 L02b
|–H. imparilis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) imparilis] L03
|–H. jinghongicus Li 1995 L02a
|–H. kobus Lee & Thornton 1967 NL07
|–H. lachlani (Enderlein 1903) [=Pseudocaecilius lachlani] SN08
|–H. leei New 1993 NL07
|–H. liangi Li 1991 L02b
|–H. loiaoi New 1991 L02a
|–H. longicornis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) longicornis] L03
|–H. longipenna Lee & Thornton 1967 NL07
|–H. longispinus Lee & Thorton 1967 NL07
|–H. maculans Li 1991 L02b
|–H. maculifornis (Thornton 1961) L02b [=Pseudocaecilius maculifrons L02a]
|–H. maculosus Lee & New 1992 NL07
|–H. marginatus Lee & New 1992 NL07
|–H. mecotrichus Li 2002 L02b
|–H. odontothelus Li 2002 L02b
|–H. orientalis Lee & New 1992 NL07
|–H. ornatus (Enderlein 1903) [=Pseudocaecilius ornatus] NL07
|–H. papillosus Vaughan, Thornton & New 1991 NL07
|–H. paulicellus (Li 1999) NL07, L02a [=*Orbocaecilius paulicellus L02a]
|–H. platokeratis Li 1993 L02a
|–H. pseudoanomalus Li 1993 L02b [=Obrocaecilius (l. c. for Orbocaecilius) pseudoanomalus L03]
|–H. quadrangulus Li 1992 L02b
|–H. reticulatus Lee & New 1992 NL07
|–H. rhadnus Thornton 1984 NL07
|–H. sedecimidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) sedecimidentus] L03
|–H. stenokeratis Li 1993 L02a
|–H. stiliger Lee & Thornton 1967 NL07
|–H. taipingensis Garcia Aldrete 1999 NL07
|–H. tartareus Li 2002 L02b
|–H. vulturius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) vulturius] L03
`–H. xihuicus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) xihuicus] L03

Chorocaecilius brunellus (Tillyard 1923) [=Caecilius brunellus, Heterocaecilius brunellus, Pseudocaecilius brunellus; incl. H. diogenes Lee & Thornton 1967, *Chorocaecilius diogenes] SN08

Pseudocaecilius citricola (Ashmead 1879) NL07 [=Psocus citricola M93; incl. Pse. elutus var. africanus Badonnel 1931 L02a, Elipsocus criniger Perkins 1899 M93, Kilauella criniger M93, Pseudocaecilius criniger M93, *Pse. elutus Enderlein 1903 NL07, L02b, Trichopsocus (*Pseudocaecilius) elutus L02a, Trichopsocus indicatus Navás 1934 M93, Pse. medianicostatus Li 1993 L02a, Caecilius pretiosus Banks 1920 M93, Pse. pretiosus M93, Pse. wolcotti Banks 1924 M93]

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l’Angola. Diamang Publicações Culturais 26: 11–267.

[L02a] Li F. 2002a. Psocoptera of China vol. 1. Science Press: Beijing.

[L02b] Li F. 2002b. Psocoptera of China vol. 2. Science Press: Beijing.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.

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