Graphipterus serrator, copyright Hectonichus.

Belongs within: Harpalinae.
Contains: Adelotopus, Sphallomorpha.

The Pseudomorphini are a group of often flattened ground beetles found living under bark. They are often found in association with ants and larvae of some species are physogastric inquilines in ant nests (Lawrence & Britton 1991). The majority of genera in this group are ovoviviparous (Baehr 2008).

Characters (from Lawrence & Britton 1991): Antennal scape not visible from above; antennae sometimes short and clavate; head with deep antennal sulcus between eye and mandible; legs more or less concealed beneath body, with may be flattened and laterally explanate.

<==Pseudomorphini [Pseudomorphinae]
|  i. s.: Cryptocephalomorpha B08
|–+–Pseudomorpha Kirby 1825 OM08, LM87
|  |    |–*P. excrucians Kirby 1825 LM87
|  |    |–P. angustata OM08
|  |    `–P. confusa Notman 1925 LM87
|  `–Graphipterus Latreille 1802 OM08, L02 [Graphipterini]
|       |–G. circumdatus Raffr. 1885 G89
|       |–G. cordiger OM08
|       |–G. limbatus OM08
|       |–G. minutus B14
|       |–G. serrator (Forskål 1775) [=Carabus serrator] G89
|       |–G. trilineatus [=Carabus trilineatus] L02
|       `–G. variegatus [=Carabus variegatus] L02
`–+–+–Adelotopus OM08
|  |–Paussotropus Waterhouse 1877 B08
|  |    `–P. cylindricus (Chaudoir 1862) (see below for synonymy) B08
|  `–Cainogenion Notman 1925 B08
|       |–*C. ipsoides (Newman 1837) [=Adelotopus ipsoides] B08
|       |    |–C. i. ipsoides B08
|       |    `–C. i. occidentale Baehr 1997 B08
|       |–C. creberrimum (Blackburn 1901) [=Adelotopus creberrimus] LM87
|       |–C. ephippiatum (Newman 1856) [=Adelotopus ephippiatus] LM87
|       `–C. obscurum (Castelnau 1867) [=Adelotopus obscurus; incl. A. subopacus Macleay 1871] LM87
`–+–Orthogonius [Orthogoniinae, Orthogoniini] OM08
|    |–O. brevilabris B55
|    |–O. clarkei B55
|    |–O. davidis B89
|    |–O. duplicatus [incl. O. alternans] B89
|    `–O. profundestriatus B89
`–Sphallomorpha OM08

Paussotropus cylindricus (Chaudoir 1862) [=Adelotopus cylindricus, Cainogenion cylindricum; incl. *Paussotropus parellelus Waterhouse 1877] B08

*Type species of generic name indicated


[B08] Baehr, M. 2008. New and rare pseudomorphine species from Western Australia (Insecta, Coleoptera, Carabidae, Pseudomorphinae). Records of the Western Australian Museum 24 (2): 151–180.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

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