Psyllipsocus ramburii, copyright Chris Lann.

Belongs within: Psocodea.

The Psocathropetae are a group of barklice with a long head and long hind legs. Wings, if present, bear a nodulus, but brachypterous and micropterous forms are common in the family Psyllipsocidae (New & Lienhard 2007).

Characters (from New & Lienhard 2007): Head long, vertical. Antennae sometimes with secondary annulations on flagellar segments. Second segment of maxillary palp without conical sensillum on inner side. Fore wing with nodulus; brachypterous individuals with some venation discernible. Scales not present on body or wings. Combined length of hind tibia and tarsus greater than abdomen. Female with gonapophyses reduced but external valve present as broad setose lobe.

<==Psocathropetae [Psocatropetae, Psyllipsocetae, Psylliposciformia]NL07
    |--Prionoglarididae [Prionoglaretae, Prionoglaridae, Prionoglaroidea]NL07
    |    |  i. s.: *Sensitibilla strinatiiNL07
    |    |--Speleketor Gurney 1943M93 [Speleketoridae, SpeleketorinaeNL07, Speleketoroidea]
    |    |    |--*S. flocki Gurney 1943M93
    |    |    |--S. irwini Mockford 1984M93
    |    |    `--S. pictus Mockford 1984M93
    |    `--Prionoglaridinae [Prionoglarinae]NL07
    |         |--*Siamoglaris zebrina Lienhard 2004NL07
    |         `--Prionoglaris Enderlein 1909 [incl. Scoliopsyllopsis Enderlein 1912]L98
    |              |--*P. stygia Enderlein 1909 [incl. Scoliopsyllopsis latreillei Enderlein 1912]L98
    |              |--P. dactyloides Lienhard 1988L98
    |              `--P. lindbergi Badonnel 1962L98
    `--Psyllipsocidae [Psyllipsocinae, Psyllipsocoidea]NL07
         |--Pseudorypteryx García Aldrete 1984M93
         |    `--*P. mexicana García Aldrete 1984M93
         |--Pseudopsyllipsocus Li 2002 [=Pseudopsyllipsous]L03
         |    `--*P. gangliigerus Li 2002 [=P. gangliigera]L03
         |--Dorypteryx Aaron 1883 [incl. Dolopteryx Smithers 1958]LL09
         |    |--*D. pallida Aaron 1883LL09 [incl. D. albicans Ribaga 1907L98]
         |    |--D. domestica (Smithers 1958) [=*Dolopteryx domestica]LL09
         |    |--D. longipennis Smithers 1991LL09
         |    `--D. yunnanica Li & Liu 2009LL09
         |--Psocathropos Ribaga 1899L02 (see below for synonymy)
         |    |--*P. lachlani Ribaga 1899NL07 (see below for synonymy)
         |    |--P. cameriferus Li 2002L02
         |    |--P. cuttackae [=*Granthakita cuttackae]NL07
         |    |--P. domesticus Li 2002L03
         |    |--P. dyadoclemus Li 2002L03
         |    |--P. pilipennisM93 [=*Gambrella pilipennisNL07]
         |    |--P. pleurotaenus Li 2002L02
         |    |--P. purpuripsarous Li 2002L02
         |    |--P. sinensis Li 1994L02
         |    |--P. termitorum (n. d.)M93 [=*Vulturops termitorumNL07]
         |    `--P. tridymus Li 2002L02
         `--Psyllipsocus Selys-Longchamps 1872L02 (see below for synonymy)
              |--*P. ramburii Selys-Longchamps 1872L98 (see below for synonymy)
              |--P. batuensis Thornton 1962NL07
              |--P. hirsutus Thornton 1962NL07
              |--P. maculatus Li 2002L02
              |--P. metamicropterus (Enderlein 1963)L02
              |--P. oculatus Gurney 1943M93
              |--P. sanxiaensis Li 1997L02
              |--P. sauteri (Enderlein 1908)L02 [=*Parempheria sauteriNL07]
              |--P. sinicus Li & Yang 1988L02
              |--P. spinosus Badonnel 1955B55
              |--P. variabilis Badonnel 1986B86
              |--P. yongi New & Lee 1992NL07
              `--P. yucatanB50

Psocathropos Ribaga 1899L02 [=PsocatroposNL07, Psocothropos (l. c.)L03; incl. Axinopsocus Enderlein 1903L98, Gambrella Enderlein 1931L98, Granthakita Behura & Dash 1968L98, Psocinella Banks 1900L98, Vulturops Townsend 1912L98; Psocatropidae]

*Psocathropos lachlani Ribaga 1899NL07 [incl. Vulturops floridensis Corbett & Hargreaves 1915L98, V. floridanus (l. c.)L98, Psocatropos lesnei Badonnel 1931M93, *Axinopsocus microps Enderlein 1903NL07, L02, Psocathropos micropsNL07, Psoquilla micropsM93, *Psocinella slossonae Banks 1900NL07, L98]

Psyllipsocus Selys-Longchamps 1872L02 [incl. Fabrella Lacroix 1915L98, Fita Navás 1913L98, Nymphopsocus Enderlein 1903L98, Ocellataria Weber 1907L98, Ocelloria Weber 1906L98, Parempheria Enderlein 1906L98]

*Psyllipsocus ramburii Selys-Longchamps 1872L98 [incl. Nymphopsocus troglodyta var. algericus Enderlein 1909L98, Psy. ramburii brachypterus Badonnel 1943L98, *Fabrella convexa Lacroix 1915NL07, L98, *Nymphopsocus destructor Enderlein 1903NL07, L98, *Ocelloria gravonymphia Weber 1906NL07, L98, *Ocellataria gravinymphaNL07, Dorypteryx hageni Banks 1897L98, Psocus pedicularius Rambur 1942 nec Hemerobius pedicularius Linnaeus 1758 nec P. pedicularius Burmeister 1839M93, Nymphopsocus troglodyta Enderlein 1909L98, Psyllipsocus (Nymphopsocus) troglodytesM93, Psy. ramburii troglodytesM93, *Fita vestigator Navás 1913NL07, L98]

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l’Angola. Diamang Publicações Culturais 26: 11–267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[B50] Broadhead, E. 1950. A revision of the genus Liposcelis Motschulsky with notes on the position of this genus in the order Corrodentia and on the variability of ten Liposcelis species. Transactions of the Royal Entomological Society of London 101 (10): 335–388, pls 1–3.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[LL09] Li, F. ,& X. Liu. 2009. Discovery in China of Dorypteryx Aaron (Psocoptera: Trogiomorpha: Psyllipsocidae), with one new species. Zootaxa 1983: 63–65.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

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