Cullen cinereum, copyright Mark Marathon.

Belongs within: Papilionoideae.

The Psoraleeae are a widespread group of legumes, mostly small trees, shrubs or subshrubs, bearing indehiscent, usually one-seeded fruits. Leaves are often dotted with glands, at least on the lower surface; members of the genus Cullen also have the fruits densely glandular. Arabian pea Bituminaria bituminosa has a distinctive strong tar-like odour. Members of the genus Psoralea are small shrubs or perennial herbs bearing solitary flowers or flowers in axillary clusters towards the ends of stems.

| |--B. bituminosaCP13 [=Psoralea bituminosaDH98]
| | |--B. b. var. bituminosaDH98
| | `--B. b. var. brachycarpa (Feldman) Danin in Danin & Hedge 1998 (see below for synonymy)DH98
| `--B. flaccidaDH98
`--+--Orbexilum lupinellumCP13
|--Otholobium bracteolatumCP13
| |--H. macrostachya [=Psoralea macrostachya]H93
| |--H. orbicularis [=Psoralea orbicularis]H93
| `--H. strobilina [=Psoralea strobilina]H93
| | |--P. californicum [=Psoralea californica]H93
| | |--P. castoreum [=Psoralea castorea]H93
| | |--P. mephiticumH93
| | `--P. pentaphyllumCP13
| `--+--Ladeania junceaCP13
| `--RupertiaCP13
| |--R. hallii [=Hoita hallii]H93
| |--R. physodes [=Psoralea physodes]H93
| `--R. rigida [=Psoralea rigida, Hoita rigida]H93
| |--P. australasicaM99
| |--P. cinereaCP13
| |--P. cuspidataJ23
| |--P. esculentaJ23
| |--P. graveolensM99
| |--P. incanaJ23
| |--P. pallidaM99
| |--P. patensM99
| |--P. pinnataB00b
| |--P. plicataD17
| `--P. tenaxG32
|--C. australasicumB00b
|--C. badoconumB00b
|--C. balsamicumB00b
|--C. cinereumB00b
|--C. cuneatumLK14
|--C. leucanthumB00a
|--C. microcephalumB00b
|--C. patensB00b
|--C. praeuptorumLK14
|--C. pustulatumB00b
|--C. tenaxCP13
`--C. virensLK14

Bituminaria bituminosa var. brachycarpa (Feldman) Danin in Danin & Hedge 1998 [=Psoralea bituminosa var. brachycarpa Feldman in Zohary 1972]DH98

*Type species of generic name indicated


[B00a] Braby, M. F. 2000a. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).

[B00b] Braby, M. F. 2000b. Butterflies of Australia: their identification, biology and distribution vol. 2. CSIRO Publishing: Collingwood (Victoria).

[CP13] Cardoso, D., R. T. Pennington, L. P. de Queiroz, J. S. Boatwright, B.-E. Van Wyk, M. F. Wojciechowski & M. Lavin. 2013. Reconstructing the deep-branching relationships of the papilionoid legumes. South African Journal of Botany 89: 58–75.

[DH98] Danin, A., & I. C. Hedge. 1998. Contributions to the flora of Jordan 2. A new species of Satureja (Labiatae) and some new records. Willdenowia 28: 135–142.

[D17] Diels, L. 1917. Beiträge zur Flora der Zentral-Sahara und ihrer Pflanzengeographie. Nach der Sammelausbeute des Freiherrn Hans Geyr von Schweppenburg. Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 54 (Beiblatt 120): 51–155.

[G32] Girault, A. A. 1932. Hymenoptera, Thysanoptera nova Australiensis. II. Privately published (reprinted: Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 292).

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[J23] James, E. 1823. Account of an Expedition from Pittsburgh to the Rocky Mountains, performed in the years 1819 and ’20, by order of the Hon. J. C. Calhoun, sec’y of war: under the command of Major Stephen H. Long. From the notes of Major Long, Mr. T. Say, and other gentlemen of the exploring party vol. 1. H. C. Carey & I. Lea: Philadelphia.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[M99] Matthews, M. 1999. Heliothine Moths of Australia: A guide to bollworms and related noctuid groups. CSIRO Publishing.

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