Amphissites alticostatus, (clockwise from top left) left, right, right interior and left interior valves, from Hoare & Merrill (2004).

Belongs within: Palaeocopida.
Contains: Primitiopsidae, Graviidae, Arcyzonidae, Scrobiculidae, Kirkbya, Polytylites.

The Kirkbyoidea are a group of reticulate, straight-backed ostracods known from the Devonian to the Middle Permian. In many examples, including members of the families Kirkbyidae, Amphissitidae and Arcyzonidae, the carapace valves are marked by a large subcentral pit (Benson et al. 1961).

| i. s.: Carnizzia auriculaL05
|--Puncioidea [Punciacea]L05
| | i. s.: PromanawaL05
| |--PunciidaeMD01
| | |--Manawa Hornibrook 1949BB61
| | | |--*M. tryphena Hornibrook 1949BB61
| | | `--M. staceyiL05
| | `--Puncia Hornibrook 1949BB61
| | `--*P. novozealandica Hornibrook 1949BB61
| |--Herrigokella [Herrigokellidae]L05
| `--TriassopunciidaeL05
|--Primitiopsoidea [Primitiopsacea, Primitiopsiomorphinina]L05
| |--PrimitiopsidaeM61a
| |--VentrigyridaeWSB93
| | |--Ventrigyrus sulcatus (Kanygin 1965)WSB93
| | `--Zenkopsis enormis (Zenkova 1975)WSB93
| |--BubnoffiopsidaeWSB93
| | |--Bubnoffiopsis bubnoffi Schrallreuter 1964WSB93
| | `--Lembitites incognitus (Sidaraviciene 1975)WSB93
| |--VenzavellidaeL05
| |--LysogorellidaeL05
| `--GraviidaeL05
`--Kirkbyoidea [Kirkbyacea, Kirkbyocopina]L05
| i. s.: ObotritiaL05
|--Cardiniferella Sohn 1953 [Cardiniferellidae]BB61
| |--*C. bowsheri Sohn 1953BB61
| `--C. ringwoodensis (Harris & Jobe 1956)WSB93
|--Kellettinidae [Kellettininae]HM04
| |--Semipetasus Sohn 1954BB61
| | `--*S. signatus Sohn 1954BB61
| |--Kindlella Sohn 1954HM04
| | |--*K. fissiloba Sohn 1954BB61
| | `--K. proscillata Hoare & Merrill 2004HM04
| `--Kellettina Swartz 1936BB61
| |--*K. robusta (Kellett 1933) [=Ulrichia robusta]BB61
| `--K. vidriensis Hamilton 1942WSB93
|--Kirkbyidae [Kirkbijidae]HM04
| |--KirkbyaHM04
| |--Villozona aspera (Polenova 1974)WSB93
| |--Carinaknightina carinata Sohn 1970WSB93
| |--Coronakirkbya Sohn 1954ZL87
| | `--*C. fimbriata Sohn 1954BB61
| |--Knightina Kellett 1933 [incl. Tenebrion Zanina 1956]BB61
| | `--*K. allorismoides (Knight 1928) [=Amphissites allorismoides]BB61
| |--Strepula Jones & Holl 1886M61b, BB61
| | `--*S. concentrica Jones & Holl 1886BB61
| `--Aurikirkbya Sohn 1950HM04
| |--*A. wordensis (Hamilton 1942) [=Kirkbya wordensis]BB61
| |--A. kellettae (Harlton 1929) [=Kirkbya kellettae, Knightina kellettae]HM04
| |--A. knighti (Harlton 1929)HM04
| `--A. triserata Shaver in Thompson et al. 1959HM04
|--Vitissites comtei Becker 1981WSB93
|--Neoamphissites costatus Becker & Wang 1992WSB93
|--Kegelites Coryell & Booth 1933HM04 (see below for synonymy)
| |--*K. spinosus (Coryell & Booth 1933) [=*Girtyites spinosus]BB61
| |--K. dattonensis (Harlton 1927) [=Amphissites dattonensis]ZL87
| |--K. harltoni (Cooper 1946) [=Ectodemites harltoni]HM04
| `--K. wapanuckensis (Harlton 1929)HM04
|--Shleesha Sohn 1961HM04
| |--S. houae (Chen 1958) [=Amphissites houae]CB86
| |--S. pinguis (Ulrich & Bassler 1906) [=Kirkbya pinguis]HM04
| |--S. pinguoides (Croneis & Gale 1939)HM04
| |--S. simplex (Roth 1929)HM04
| `--S. sullivanensis (Payne 1937)HM04 [=Shleesha sullivanensisZL87, Ectodemites sullivanensisZL87]
`--Amphissites Girty 1910HM04 (see below for synonymy)
| i. s.: A. centronotus (Ulrich & Bassler 1906)ZL87 (see below for synonymy)
| A. primus [=*Ectodemites primus]BB61
| A. robertsi Morey 1935ZL87
`--A. (Amphissites)HM04
|--*A. (A.) rugosus Girty 1910BB61
|--A. (A.) alticostatus Bradfield 1935HM05
|--A. (A.) insignis Croneis & Thurman 1939HM04
|--A. (A.) morrowanensis Hoare & Merrill 2004HM04
`--A. (A.) reticulatus Geis 1932HM04

Amphissites Girty 1910HM04 [incl. Albanella Harris & Lalicker 1932BB61, Binodella Bradfield 1935BB61, Brillius Brayer 1952BB61, Ectodemites Cooper 1941BB61]

Amphissites centronotus (Ulrich & Bassler 1906)ZL87 [=Kirkbya centronotaZL87; incl. A. blumenstengeli Gründel 1962S82, A. mosquensis Posner 1951S82]

Kegelites Coryell & Booth 1933HM04 [=Girtyites Coryell & Booth 1933 non Wedekind 1914BB61; incl. Kirkbyites Johnson 1936BB61]

*Type species of generic name indicated


[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[CB86] Chen D.-Q. & Bao H. 1986. Lower Permian ostracodes from the Chihsia Formation of Jurong and Longtan, Jiangsu Province. Acta Micropalaeontologica Sinica 3 (2): 107–132.

[HM04] Hoare, R. D., & G. K. Merrill. 2004. A Pennsylvanian (Morrowan) ostracode fauna from Texas. Journal of Paleontology 78 (1): 185–204.

[L05] Liebau, A. 2005. A revised classification of the higher taxa of the Ostracoda (Crustacea). Hydrobiologia 538: 115–137.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[M61a] Moore, R. C. 1961a. Summary of classification and stratigraphic distribution. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q92–Q99. Geological Society of America and University of Kansas Press.

[M61b] Moore, R. C. 1961b. Supplement—USSR treatise on Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q422–Q429. Geological Society of America and University of Kansas Press.

[S82] Shi C.-G. 1982. Some early Carboniferous ostracodes from Nylam, Xizang (Tibet). Acta Palaeontologica Sinica 21 (3): 309–314.

[WSB93] Whatley, R. C., D. J. Siveter & I. D. Boomer. 1993. Arthropoda (Crustacea: Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 343–356. Chapman & Hall: London.

[ZL87] Zhang X.-J. & Liang X.-Y. 1987. Ostracoda from the Taiyuan Formation of Xingyang and Gong Xian districts, Henan. Acta Micropalaeontologica Sinica 4 (3): 293–312.

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