Pyrgulina pseudorugata, from Marshall & Murdoch (1921).

Belongs within: Pylopulmonata.
Contains: Syrnola, Pyramidella, Strioturbonilla, Turbonilla, Chemnitzia, Eulimella, Menestho, Chrysallida, Odostomia, Evalea.

The Pyramidellidae are a group of mostly minute marine snails with a heterostrophic protoconch and no anterior canal on the shell aperture. Members of the family lack a radula and live as ectoparasites of other invertebrates such as other molluscs or polychaetes.

|–Syrnolinae [Syrnolidae] BR05
|    |–Syrnola BR05
|    `–Tiberia Jeffreys 1884 [Tiberiinae, Tiberiini] BR05
|         |–*T. minuscula (Monterosato 1880) [=Pyramidella minuscula] BR17
|         `–T. nitidula (Adams 1860) [=Syrnola nitidula] BR17
|–Pyramidellinae BR05
|    |–Sayella Dall 1885 [Sayellinae, Sayellini] BR05
|    |    |–*S. hemphillii (Dall 1883) [=Leuconia hemphillii] BR17
|    |    `–S. crosseana Dall 1885 MP65
|    `–Pyramidellini BR05
|         |–Pyramidella BR05
|         `–Plotia Röding 1798 [Plotiidae] BR05
|              |–*P. lineata Röding 1798 BR17
|              `–P. scabra [=Melania (Plotia) scabra; incl. M. denticulata Lea 1851, M. doreyana, M. spinulosa] TC89
|–Turbonillinae [Chemnitziidae, Chemnitziinae, Turbonillidae] BR05
|    |  i. s.: Strioturbonilla O27a
|    |–Turbonillini BR17
|    |    |–Turbonilla BR05
|    |    `–Chemnitzia BR17
|    |–Cingulina Adams 1860 BR05 [Cingulininae, Cingulinini BR17]
|    |    |–*C. circinata (Adams 1860) BR17
|    |    `–C. isseli (Tryon 1886) [incl. Eulimella cingulata Issel 1869, Odetta cingulata] BD86
|    `–Eulimellini BR17
|         |–Eulimella BR05
|         `–Belonidium Cossmann 1893 non Mont. & Durieu 1846 (ICBN) BR17 [=Anisocycla Monterosato 1880 BR05]
|              `–*B. gracile (Deshayes 1861) [=Aciculina gracilis, *Anisocycla gracilis] BR17
`–Odostomiinae [Odostomiidae] BR05
|–Odostomella Bucquoy, Dautzenberg & Dollfus 1883 [Odostomellidae, Odostomellinae, Odostomellini] BR05
|    `–*O. doliolum (Philippi 1844) [=Rissoa doliolum] BR17
|–Cyclostremella Bush 1897 [Cyclostremellidae, Cyclostremellinae, Cyclostremellini] BR05
|    |–*C. humilis Bush 1897 BR17
|    |–C. californica Bartsch 1907 O27b
|    `–C. concordia Bartsch 1920 O27b
|–Chrysallidini [Chrysallididae, Chrysallidinae] BR05
|    |–Menestho BR05
|    |–Chrysallida BR05
|    `–Pyrgulina Adams 1864 [Pyrgulininae] BR05
|         |–*P. casta (Adams 1861) [=Chrysallida casta] BR17
|         |–P. gliriella Melvill & Standen 1896 H09
|         |–P. henni (Brazier 1894) [=Odostomia henni] H09
|         |–P. pseudorugata M. & M. 1921 F27
|         |–P. rugata F27
|         |–P. senex Hedley 1902 H09
|         |–P. umeralis Hedley 1902 H09
|         `–P. zea Hedley 1902 H09
`–Odostomiini [Liostomiini, Liostomini] BR05
|–Odostomia BR05
|–Spiralinella pellucida (Dillwyn 1817) BR05
|–Evalea O27a
|–Liostomia Sars 1878 BR05
|    `–*L. clavula (Lovén 1846) [=Turbonilla clavula] BR17
|–Ividella Dall & Bartsch 1909 O27a
|    |–*I. navisa (Dall & Bartsch 1907) P61, O27a [=Odostomia (*Ividella) navisa P61]
|    |    |–I. n. navisa O27a
|    |    `–I. n. delmontensis (Dall & Bartsch 1907) [=Odostomia (Ividella) navisa delmontensis] O27a
|    |–I. maoria Powell 1940 P61
|    `–I. pedroana (Dall & Bartsch 1909) [=Odostomia (Ividella) pedroana] O27a
|–Miralda Adams 1864 O27a
|    |–M. aepynota (Dall & Bartsch 1909) [=Odostomia (Miralda) aepynota] O27a
|    |–M. hemphilli (Dall & Bartsch 1909) [=Odostomia (Miralda) hemphilli] O27a
|    |–‘Parthenia’ quinquecincta [incl. Rissoa notabilis, Cingula turrita] C64
|    |–M. soteloi Rolán 1994 BC01
|    |–M. superba Rolán & Fernandes 1993 BC01
|    `–M. temperata Rolán & Fernandes 1993 BC01
`–Besla Dall & Bartsch 1904 O27a, P61
|–*B. convexa [=Chrysallida convexa] P61
|–B. callimorpha (Dall & Bartsch 1909) (see below for synonymy) O27a
|–B. rossiana Laws 1941 P61
|–B. subrugata (Powell 1927) P61
|–B. vaga Laws 1941 P61
`–B. waitangiensis Laws 1941 P61

Pyramidellidae incertae sedis:
Myxa Hedley 1903 F27
Kleinella fulva (Adams 1851) [=Monoptygma fulva, Pyrammidella (Actaeopyramis) fulva] BD86
Monoptygma sinuata Gould 1861 BD86
Otopleura mitralis Adams 1855 BW09
Pseudoscilla verdensis Peñas & Rolán 1999 BC01
Agatha Adams 1860 P61
|–*A. virgo [=Pyramidella virgo] P61
`–A. georgiana (Hutton 1885) P61 (see below for synonymy)
Obexomia Laws 1941 [incl. Obex Laws 1940] P61
`–*O. denselirata (Suter 1908) [=Odostomia denselirata] P61
Striodostomia Laws 1940 P61
`–*S. orewa Laws 1940 P61
Gumina Finlay 1928 P61
|–*G. dolichostoma (Suter 1908) [=Odostomia dolichostoma] P61
`–G. minor Laws 1940 P61
Linopyrga Laws 1941 P61
`–*L. rugata (Hutton 1886) [=Odostomia rugata] P61
|–L. r. rugata P61
`–L. r. parva Laws 1941 P61
Siogamaia Nomura 1936 P61
|–*S. fortiplicata [=Tropaeas (*Siogamaia) fortiplicata] P61
`–S. morioria Laws 1941 P61
Puposyrnola Cossmann 1921 P61
|–*P. acicula [=Auricula acicula] P61
|–P. fastigiata (Suter 1906) P61
`–P. missile Laws 1937 P61
Terelimella Laws 1937 P61
|–*T. ototarana P61
|–T. aupouria (Powell 1937) P61
|–T. benthicola Dell 1956 P61
`–T. larochei (Powell 1930) P61
Finlayola Laws 1937 P61
|–*F. finlayi P61
`–F. lurida (Suter 1908) P61
Planpyrgiscus Laws 1937 P61
|–*P. extenuatus [=Turbonilla extenuata] P61
`–P. lawsi Dell 1956 P61
Pyrgiscilla Laws 1937 P61
|–*P. chattonensis [=Turbonilla chattonensis] P61
`–P. otakauica Laws 1937 P61
Striarcana Laws 1937 P61
|–*S. cryptolira Laws 1937 P61
`–S. tauranga Laws 1937 P61
Creonella TTE93
|–C. subangulata Sohl 1964 TTE93
|–C. triplicata Wade 1917 TTE93
`–C. turretiforma Sohl 1964 TTE93

Agatha georgiana (Hutton 1885) P61 [=Odostomia georgiana F27; incl. O. bembix Suter 1908 P61, F27, O. inornata P61, O. stygia P61]

Besla callimorpha (Dall & Bartsch 1909) [=Odostomia (Besla) callimorpha, Chrysallida pumila Carpenter 1866 non O. pumila Adams 1861] O27a

*Type species of generic name indicated


[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177–208.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[MP65] Merrill, A. S., & R. E. Petit. 1965. Mollusks new to South Carolina. Nautilus 79 (2): 58–66.

[O27a] Oldroyd, I. S. 1927a. The Marine Shells of the West Coast of North America vol. 2 pt 2. Stanford University Press: Stanford University (California).

[O27b] Oldroyd, I. S. 1927b. The Marine Shells of the West Coast of North America vol. 2 pt 3. Stanford University Press: Stanford University (California).

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[TC89] Tapparone Canefri, T. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XVIII.—Molluschi terrestri e d’acqua dolce. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 295–359.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

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