Racomitrioideae

Bucklandiella microcarpa, copyright Nataliya Gamova.

Belongs within: Bryopsida.
Contains: Racomitrium section Canescentia, Bucklandiella section Laevifoliae.

Bucklandiella lusitanica
Published 29 September 2020

The diversity of mosses is much higher than many people realise. Whereas some moss species have wide ranges that may cross between continents and hemispheres, others are unique to very specific regions and habitats. Among examples of the latter is the European species Bucklandiella lusitanica.

Illustrations of Bucklandiella lusitanica, from Ochyra & Sérgio (1992). Top left: habit; top right: section of stem of hair-leafed form when dry; lower left: section of stem of hairless form and sporophyte when wet.

Bucklandiella lusitanica was only described as a new species (under the name Racomitrium lusitanicum) in 1992 (Ochyra & Sérgio 1992), having gone unnoticed previously despite being a relatively distinctive species. Recent collections of the species have been identified from a single region, the Serra do Gerês mountain rainge and Parque Natural da Peneda-Gerês national park in the northwest of Portugal, at altitudes between 650 and 1000 metres. A single collection from the Serra do Estrela further south in the country was made in the mid-1800s though it went unidentified at the time. Its rarity is such that is has officially been listed as Endangered by the IUCN. Bucklandiella lusitanica is a rheophyte, which is to say that it grows in association with running water. It grows on acidic granite rocks that are periodically or permanently submerged, such as alongside streams and waterfalls. It is particularly abundant on steep rock faces, growing in association with closely related moss species.

Appearance-wise, Bucklandiella lusitanica is a medium-sized moss with irregularly branched stems growing 1.5 to 3.5 centimetres in length. Leaves are rigid, held tight to stem, and two or three millimetres long.One of the species’ most distinctive features is a broad, fleshy margin to each leaf that is generally two or three cells thick whereas the lamina of the leaf is mostly only a single cell thick. The alar cells at the base of the sides of the leaf often form inflated, strongly coloured lobes. The leaves commonly end in a fine, colourless hair-point. The structure of the leaves is similar to that of Bucklandiella lamprocarpa, another aquatic moss species, but that species lacks the hair-points. The two species also differ in the form of their spores, those of B. lamprocarpa being larger and more ornate than those of B. lusitanica, and B. lamprocarpa has fatter and often shinier capsules than B. lusitanica.

I mentioned previously that Bucklandiella lusitanica was originally described as a member of the genus Racomitrium. The moss genus Racomitrium was long recognised by a distinctive array of features including leaf lamina cells with distinctly sinuous longitudinal cell walls, a calyptra (the cap of the developing capsule) that is basally frayed into several lobes, and teeth of the peristome (the teeth around the aperture of a mature capsule) that are split into two or more segments (Sawicki et al. 2015). Racomitrium in this sense was a diverse genus with over two hundred species having been named at one time or another, and somewhere between sixty and eighty species recognised as valid in recent years, As a result, Ochyra et al. (2003) proposed the division of Racomitrium in the broad sense between four separate genera. Bucklandiella, the largest of these segregate genera (with about fifty currently known species), was recognised for species with a smooth leaf surface (lacking papillae on the lamina) and relatively short, shallowly divided teeth in the peristome. The division of Racomitrium has not been universally accepted. Larrain et al. (2013) questioned the monophyly and diagnosability of Ochyra et al.‘s segregates but Sawicki et al. (2015) reiterated their support for the new system (and added a fifh new segregate genus to boot). It is generally accepted that Racomitrium in the broad sense represents a monophyletic unit, so the question of whether lusitanicum should be assigned to Racomitrium or Bucklandiella may largely be considered a question of just how closely circumscribed you feel a genus should be.

Systematics of Racomitrioideae
RacomitrioideaeOZB-O03
|--Codriophorus P.Beauv. 1822OZB-O03
| |--*C. acicularis (Hedw.) Beauv. 1823OZB-O03 (see below for synonymy)
| `--C. corrugatusB-OO10
|--Racomitrium Brid. 1819S04 (see below for synonymy)
| | i. s.: R. aduncoides Bednarek-Ochyra 1999 (see below for synonymy)OZB-O03
| | R. anomodontoides Cardot 1908 (see below for synonymy)OZB-O03
| | R. brevisetum Lindb. 1872 (see below for synonymy)OZB-O03
| | R. carinatum Cardot 1908 (see below for synonymy)OZB-O03
| | R. crispulum (Hooker & Wilson) Hooker & Wilson 1854 [=Grimmia crispula (preoc.)]SK02
| | |--R. c. var. crispulum (see below for synonymy)SK02
| | `--R. c. var. tasmanicum (Hampe) Lawton 1973 [=R. heterostichum var. tasmanicum]SK02
| | R. depressum Lesq. 1868 (see below for synonymy)OZB-O03
| | R. dichelymoides Herzog 1934 (see below for synonymy)OZB-O03
| | R. hespericum Sérgio, Muñoz & Ochyra 1995 (see below for synonymy)OZB-O03
| | R. laevigatum Jaeger 1874 (see below for synonymy)OZB-O03
| | R. molle Cardot 1908 (see below for synonymy)OZB-O03
| | R. nigritumD03
| | R. norrisii Bednarek-Ochyra & Ochyra 2000 (see below for synonymy)OZB-O03
| | R. papeetense Besch. 1894 (see below for synonymy)OZB-O03
| | R. pseudopatens (Müll.Hal.) Paris 1900 [=Grimmia pseudopatens]SK02
| | R. ptychophyllum (Mitt.) Mitt. 1867SK02
| | R. rupestre (Hooker & Wilson) Hooker & Wilson 1854 [=R. crispulum var. rupestre]SK02
| | R. ryszardii Bednarek-Ochyra 2000 (see below for synonymy)OZB-O03
| | R. sublamprocarpumD03
| | R. subsecundumN02
| | ‘Grimmia’ varia Mitt. 1864 (see below for synonymy)OZB-O03
| |--R. sect. Racomitrium [incl. R. [unranked] Lanuginosa Kindb. 1898, R. sect. Lanuginosa (Kindb.) Nog. 1974]OZB-O03
| | |--*R. lanuginosum (Hedw.) Brid. 1819OZB-O03, S04 (see below for synonymy)
| | | |--R. l. ssp. lanuginosumOZB-O03
| | | `--R. l. ssp. geronticumOZB-O03
| | `--R. patagonicumOZB-O03
| |--R. sect. CanescentiaS04
| `--R. sect. Stenotrichum (Chev.) Bedn.-Ochyra 1995S04 [incl. R. subg. Cataracta Vilhelm 1926OZB-O03]
| |--R. aquaticum (Brid. ex Schrad.) Brid. 1819S04 (see below for synonymy)
| `--R. fasciculare (Hedw.) Brid. 1819S04 (see below for synonymy)
`--Bucklandiella Roiv. 1972 (see below for synonymy)OZB-O03
| i. s.: *B. bartramii (Roiv.) Roiv. 1972 [=*Bucklandia bartramii Roiv. 1955]OZB-O03
| B. afoninae (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. albipilifera (Gao & Cao) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| |--B. a. var. albipiliferumOZB-O03
| `--B. a. var. lorifolia (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. angustifolia (Broth.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. brevipes (Kindb.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. compactula (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. crispipila (Taylor) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. crispula (Hook.f. & Wilson) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. cucullata (Broth.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. cucullatifolia (Hampe) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. cylindropyxis (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. decurrens (Dixon) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. defoliata (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. didymaB-OO10
| B. emersa (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 (see below for synonymy)
| B. fuscescens (Wilson) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. fuscolutea (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. gracillima (Dixon) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. grimmioides (Herzog) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. heterostichoides (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. himalayana (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. joseph-hookeri (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. laeta (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. lamprocarpa (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. lawtonae (Ireland) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. lepervanchei (Besch.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. lusitanica (Ochyra & Sérgio) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. membranacea (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. minuta (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. nitidula (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. obesa (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. occidentalis (Renauld & Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. ochracea (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. orthotrichacea (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. pachydictyon (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. pacifica (Ireland & Spence) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. procumbens (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 (see below for synonymy)
| B. ptychophylla (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. rupestris (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. seychellara (Besch.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. striatipila (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 (see below for synonymy)
| B. suborthotrichacea (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. subsecunda (Hook. & Grev.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. subulifolia (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. valdon-smithii (Ochyra & Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. venusta (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. verrucosa (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| |--B. v. var. verrucosaOZB-O03
| `--B. v. var. emodensis (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. visnadiae (Buck) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. vulcanicola (Frisvoll & Deguchi) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. subsect. Cucullaria (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. subsect. Diaphanae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. subsect. Grimmiaeformes (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| B. subsect. Horridae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
|--B. sect. Emersae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
|--B. sect. LaevifoliaeB-OO10
|--B. sect. Lawtoniae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
|--B. sect. Marginatae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
| `--B. microcarpa (Hedw.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 (see below for synonymy)
| |--B. m. f. microcarpaOZB-O03
| `--B. m. f. afoninae (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
|--B. sect. Ptychophyllae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
`--B. sect. Subsecundae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03
`--B. subsect. Shevockiella Bednarek-Ochyra & Ochyra 2010B-OO10
`--B. (subsect. *Shevockiella) shevockii Bednarek-Ochyra & Ochyra 2010B-OO10

Nomina nuda: Racomitrium austrosudeticum Watts & Whitel. 1902SK02
Rhacomitrium flavopallidum Dusén 1903D03

Bucklandiella Roiv. 1972 [=Bucklandia Roiv. 1955 non Sternb. 1825; incl. R. subg. Microcarpa Vilh. 1926, R. subg. Microcarpae]OZB-O03

Bucklandiella subsect. Cucullaria (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium subsect. Cucullaria Bednarek-Ochyra 1995]OZB-O03

Bucklandiella subsect. Diaphanae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium subsect. Diaphana Bednarek-Ochyra & Ochyra 1996]OZB-O03

Bucklandiella sect. Emersae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium sect. Emersa Bednarek-Ochyra 1995]OZB-O03

Bucklandiella subsect. Grimmiaeformes (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium subsect. Grimmiiformia Bednarek-Ochyra 1995]OZB-O03

Bucklandiella subsect. Horridae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium subsect. Horrida Bednarek-Ochyra & Ochyra 1996]OZB-O03

Bucklandiella sect. Lawtoniae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium sect. Lawtonia Bednarek-Ochyra 1995]OZB-O03

Bucklandiella sect. Marginatae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium sect. Marginata Bednarek-Ochyra 1995]OZB-O03

Bucklandiella sect. Ptychophyllae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium sect. Ptychophylla Bednarek-Ochyra 1995]OZB-O03

Bucklandiella sect. Subsecundae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium sect. Subsecunda Bednarek-Ochyra 1995]OZB-O03

Bucklandiella sect. Sudeticae (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium sect. Sudetica Bednarek-Ochyra 1995]OZB-O03

Bucklandiella afoninae (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium afoninae Frisvoll 1988]OZB-O03

Bucklandiella albipilifera (Gao & Cao) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium albipiliferum Gao & Cao in Gao, Zhang & Cao 1981]OZB-O03

Bucklandiella albipilifera var. lorifolia (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium capillifolium var. lorifolium Frisvoll 1988]OZB-O03

Bucklandiella angustifolia (Broth.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium angustifolium Broth. in Hand.-Mazzetti 1929]OZB-O03

Bucklandiella brevipes (Kindb.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium brevipes (Kindb. in Macoun 1890]OZB-O03

Bucklandiella compactula (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia compactula Müll.Hal. 1898]OZB-O03

Bucklandiella crispipila (Taylor) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Trichostomum crispipilum Taylor 1846]OZB-O03

Bucklandiella crispula (Hook.f. & Wilson) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Didimodon crispulus Hook.f. & Wilson 1844]OZB-O03

Bucklandiella cucullata (Broth.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium cucullatum Broth. 1929]OZB-O03

Bucklandiella cucullatifolia (Hampe) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium cucullatifolium Hampe 1863]OZB-O03

Bucklandiella cylindropyxis (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia cylindropyxis Müll.Hal. 1898]OZB-O03

Bucklandiella decurrens (Dixon) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium decurrens Dixon in Christoph.) 1960]OZB-O03

Bucklandiella defoliata (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia defoliata Müll.Hal. 1883]OZB-O03

Bucklandiella emersa (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 [=Grimmia emersa Müll.Hal. 1851OZB-O03, Racomitrium emersumSK02]

Bucklandiella fuscescens (Wilson) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium fuscescens Wilson in Mitt. & Wilson 1857]OZB-O03

Bucklandiella fuscolutea (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium fuscolutea Cardot 1916]OZB-O03

Bucklandiella gracillima (Dixon) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium gracillimum Dixon in Christoph. 1960]OZB-O03

Bucklandiella grimmioides (Herzog) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium grimmioides Herzog 1957]OZB-O03

Bucklandiella heterostichoides (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium heterostichoides Cardot 1905]OZB-O03

Bucklandiella himalayana (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia himalayana Mitt. 1859]OZB-O03

Bucklandiella joseph-hookeri (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium joseph-hookeri Frisvoll 1988]OZB-O03

Bucklandiella laeta (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium laetum Besch. & Cardot 1908]OZB-O03

Bucklandiella lamprocarpa (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia lamprocarpa Müll.Hal. 1849]OZB-O03

Bucklandiella lawtonae (Ireland) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium lawtonae Ireland 1970]OZB-O03

Bucklandiella lepervanchei (Besch.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium lepervanchei Besch. 1880]OZB-O03

Bucklandiella lusitanica (Ochyra & Sérgio) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium lusitanicum Ochyra & Sérgio 1992]OZB-O03

Bucklandiella membranacea (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia membranacea Mitt. 1876]OZB-O03

Bucklandiella microcarpa (Hedw.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 [=Trichostomum microcarpon Hedw. 1801OZB-O03, Grimmia microcarpaD24, Racomitrium microcarponOZB-O03; incl. G. ramulosaD24, R. ramulosumOZB-O03]

Bucklandiella microcarpa f. afoninae (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 non B. afoninae (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium microcarpon f. afoninae Frisvoll 1988 non R. afoninae Frisvoll 1988]OZB-O03

Bucklandiella minuta (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia minuta Müll.Hal. 1883]OZB-O03

Bucklandiella nitidula (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium nitidulum Cardot 1908]OZB-O03

Bucklandiella obesa (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium obesum Frisvolll 1988]OZB-O03

Bucklandiella occidentalis (Renauld & Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium heterostichum var. occidentale Renauld & Cadot 1890]OZB-O03

Bucklandiella ochracea (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia ochracea Müll.Hal. 1883]OZB-O03

Bucklandiella orthotrichacea (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia orthotrichacea Müll.Hal. 1883]OZB-O03

Bucklandiella pachydictyon (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium pachydictyon Cardot 1908]OZB-O03

Bucklandiella pacifica (Ireland & Spence) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium pacificum Ireland & Spence 1987]OZB-O03

Bucklandiella procumbens (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 [=Grimmia procumbens Mitt. 1882OZB-O03, Guembelia procumbens (n. n.)SK02]

Bucklandiella ptychophylla (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia ptychophylla Mitt. in Lindsay 1866]OZB-O03

Bucklandiella rupestris (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Dryptodon rupestris Hook.f. & Wilson 1844]OZB-O03

Bucklandiella seychellara (Besch.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium seychellarum Besch. 1880]OZB-O03

Bucklandiella striatipila (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 [=Racomitrium striatipilum Cardot 1905OZB-O03; incl. R. crispipulumSK02]

Bucklandiella suborthotrichacea (Müll.Hal.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Grimmia suborthotrichacea Müll.Hal. 1883]OZB-O03

Bucklandiella subsecunda (Hook. & Grev.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Trichostomum subsecundum Hook. & Grev. 1836]OZB-O03

Bucklandiella subulifolia (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium subulifolium Cardot 1908]OZB-O03

Bucklandiella valdon-smithii (Ochyra & Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium valdon-smithii Ochyra & Bednarek-Ochyra 1999]OZB-O03

Bucklandiella venusta (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium venustum Frisvoll 1988]OZB-O03

Bucklandiella verrucosa (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium verrucosum Frisvoll 1988]OZB-O03

Bucklandiella verrucosa var. emodensis (Frisvoll) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium verrucosum var. emodense Frisvoll 1988]OZB-O03

Bucklandiella visnadiae (Buck) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium visnadiae Buck 1999]OZB-O03

Bucklandiella vulcanicola (Frisvoll & Deguchi) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Racomitrium vulcanicola Frisvoll & Deguchi 1988]OZB-O03

*Codriophorus acicularis (Hedw.) Beauv. 1823OZB-O03 [=Bryum aciculareD24, Dicranum aciculareOZB-O03, Grimmia acicularisD24, Racomitrium aciculare (Hedw.) Brid. 1819S04, Trichostomum aciculareOZB-O03; incl. Rhacomitrium aciculare var. denticulatumD24]

‘Grimmia’ varia Mitt. 1864 [=Codriophorus varius (Mitt.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium Brid. 1819S04 [=RhacomitriumS04; incl. Ra. subsect. Andicola Bednarek-Ochyra 1995OZB-O03, Codriophorus subsect. Andicola (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03, Ra. sect. Chrysea Bednarek-Ochyra 1995OZB-O03, C. sect. Chrysei (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03, Ra. sect. Fascicularia Bednarek-Ochyra 1995OZB-O03, C. sect. Fascicularia (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03, Ra. subsect. Hydrophilum Bednarek-Ochyra 1995OZB-O03, C. subsect. Hydrophilus (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03, Ra. sect. Papillosa Kindberg 1898OZB-O03, Ra. sect. Pilifera Bednarek-Ochyra 1995OZB-O03, C. sect. Piliferi (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03]

Racomitrium aduncoides Bednarek-Ochyra 1999 [=Codriophorus aduncoides (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium anomodontoides Cardot 1908 [=Codriophorus anomodontoides (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium aquaticum (Brid. ex Schrad.) Brid. 1819S04 [=Trichostomum aquaticum Brid. ex Schrad. 1803OZB-O03, Codriophorus aquaticus (Brid. ex Schrad.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03, Racomitrium (subg. *Cataracta) aquaticumOZB-O03; incl. R. cataractarumOZB-O03, R. protensumS04]

Racomitrium brevisetum Lindb. 1872 [=Codriophorus brevisetus (Lindb.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium carinatum Cardot 1908 [=Codriophorus carinatus (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium crispulum (Hooker & Wilson) Hooker & Wilson 1854 var. crispulum [incl. R. chlorocarpum, Dryptodon chlorocarpus (n. n.), Grimmia chlorocarpa (n. n.), G. sullivanii, R. sullivanii, R. symphyodon, Grimmia symphyodonta, R. symphyodontum]SK02

Racomitrium depressum Lesq. 1868 [=Codriophorus depressus (Lesq.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium dichelymoides Herzog 1934 [=Codriophorus dichelymoides (Herzog) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium fasciculare (Hedw.) Brid. 1819S04 [=Codriophorus fascicularis (Hedw.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03, Grimmia fascicularisD24, Trichostomum fasciculare Hedw. 1801OZB-O03]

Racomitrium hespericum Sérgio, Muñoz & Ochyra 1995 [=Codriophorus hespericus (Sérgio, Muñoz & Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium laevigatum Jaeger 1874 [=Codriophorus laevigatus (Jaeger) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

*Racomitrium lanuginosum (Hedw.) Brid. 1819OZB-O03, S04 [=Grimmia lanuginosaSK02, Trichostomum lanuginosumD24; incl. Bryum hypnoides (nom. illeg.)SK02, Grimmia hypnoidesD24, Racomitrium (*Lanuginosa) hypnoides (nom. illeg.)OZB-O03, R. sundaicumSK02, Grimmia sundaicaSK02]

Racomitrium molle Cardot 1908 [=Codriophorus mollis (Cardot) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium norrisii Bednarek-Ochyra & Ochyra 2000 [=Codriophorus norrisii (Bednarek-Ochyra & Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium papeetense Besch. 1894 [=Codriophorus papeetense (Besch.) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

Racomitrium ryszardii Bednarek-Ochyra 2000 [=Codriophorus ryszardii (Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003]OZB-O03

*Type species of generic name indicated

References

[B-OO10] Bednarek-Ochyra, H., & R. Ochyra. 2010. Bucklandiella shevockii (Bryophyta, Grimmiaceae), an exquisite new species from Yunnan, China. Polish Botanical Journal 55 (2): 499–506.

[D24] Dixon, H. N. 1924. The Student’s Handbook of British Mosses 3rd ed. V. V. Sumfield: Eastbourne.

[D03] Dusén, P. 1903. Patagonian and Fuegian mosses. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 1896–1899 vol. 8. Botany pp. 63–126. The University: Princeton (New Jersey).

Larraín, J., D. Quandt, M. Stech & J. Muñoz. 2013. Lumping or splitting? The case of Racomitrium (Bryophytina: Grimmiaceae). Taxon 62 (6): 1117–1132.

[N02] Negi, H. R. 2002. Abundance and diversity of moss communities of Chopta-Tunganath in the Garhwal Himalaya. Journal of the Bombay Natural History Society 99 (3): 418–433.

Ochyra, R., & C. Sérgio. 1992. Racomitrium lusitanicum (Musci, Grimmiaceae), a new species from Europe. Fragmenta Floristica et Geobotanica 37 (1): 261–271.

[OZB-O03] Ochyra, R., J. Żarnowiec & H. Bednarek-Ochyra. 2003. Census Catalogue of Polish Mosses. Institute of Botany, Polish Academy of Sciences: Cracow.

Sawicki, J., M. Szczecińska, H. Bednarek-Ochyra & R. Ochyra. 2015. Mitochondrial phylogenomics supports splitting the traditionally conceived genus Racomitrium (Bryophyta: Grimmiaceae). Nova Hedwigia 100 (3–4): 293–317.

[S04] Smith, A. J. E. 2004. The Moss Flora of Britain and Ireland 2nd ed. Cambridge University Press.

[SK02] Streimann, H., & N. Klazenga. 2002. Catalogue of Australian Mosses. Flora of Australia Supplementary Series 17. Australian Biological Resources Study: Canberra.

Leave a comment

Your email address will not be published. Required fields are marked *