Daphnella rissoides, copyright G. & Ph. Poppe.

Belongs within: Conoidea.
Contains: Asperdaphne (Asperdaphne), Xanthodaphne, Pseudodaphnella, Eubela, Rugobela, Buccinaria, Nepotilla, Philbertia, Hemilienardia, Raphitoma, Veprecula, Eucyclotoma, Kermia, Pleurotomella (Pleurotomella), Taranis, Gymnobela, Daphnella (Daphnella).

The Raphitomidae is a very diverse group of conoid gastropods, many species of which show a reduction in the radular apparatus or other structures of the foregut (Bouchet et al. 2011). Members include the distinctive West Pacific genus Thatcheria in which the shell has a pagodiform spire with a sharp carina above middle whorl height below a broad flat shoulder. Species of the genus Daphnella have fairly smooth or delicately lirate shells with a short anterior canal and weak or reversed L-shaped sinus (Powell 1966).

Characters (from Bouchet et al. 2011): Shell very variable in shape and size, from buccinoid to ovate or to elongate-fusiform or high-cylindrical and from about 2 to over 140 mm high. Sculpture variously developed, from nearly smooth shell to well-developed spiral and axial elements; subsutural ramp, when separated from rest of the whorl, usually smooth or with traces of anal sinus growth. Apertural armature rarely well expressed, inner lip usually smooth. Anal sinus either subsutural, shaped as a ‘reversed-L’, or on subsutural ramp, asymmetric, very shallow to rather deep, typically not constrained but sometimes even almost tubular. Siphon short to rather long. Protoconch typically planktotrophic, multispiral, of 2.5–6.5 whorls, protoconch I often spirally striated, protoconch II with diagonally cancellated sculpture. Paucispiral protoconch with predominantly spiral striation. Operculum absent. Radula of hypodermic marginal teeth of very variable morphology. Teeth relatively very long to very short and reduced. Solid base of teeth from very small to rather large, side projections around base sometimes present. Barb(s) present or absent; cockscomb-shaped structure present near tooth tip in at least one species. Tooth canal opening laterally or subapically. Reduction in radular apparatus and other structures of anterior foregut frequent, grading to complete absence.

<==Raphitomidae [Daphnellinae, Daphnellini, Pleurotomellinae, Raphitominae, Thatcheriidae, Thatcheriinae]
    |  `--RaphitomaPK11
       |  `--Rimosodaphnella Cossmann 1915PK11, BK11 [=Rimosodalphnella (l. c.)P66]
       |       |--*R. textile (Brocchi 1814) [=Murex textilis]P66
       |       |--R. ciprianii (Hornung 1920) [=Daphnella (Rimosodaphnella) ciprianii]P66
       |       `--R. salinasi (Calcara 1841) [=Pleurotoma salinasi]P66
       `--+--Glyphostomoides Shuto 1983PK11, BK11
          |--Tritonoturris Dall 1924PK11, BK11
          |    |--*T. robillardi (Adams 1869) [=Clathurella robillardi]P66
          |    |--T. amabilis (Hinds 1843)P66 [=Clavatula amabilisP66, Clathurella amabilisH09]
          |    |--T. capensis (Smith 1882) [=Pleurotoma (Defrancia) capensis]P66
          |    |--T. cumingii (Powys in Powys & Sowerby 1835)BK11 [=Buccinum cumingiiP66]
          |    |--T. elegans (Pease 1860)P66 (see below for synonymy)
          |    |--T. menecharmes (Melvill 1923)P66 (see below for synonymy)
          |    `--T. ‘paucicostata’ (Adams 1869) [=Clathurella paucicostata non Pease 1860]P66
             |  `--KermiaPK11
             `--+--Pleurotomella Verrill 1873PK11, BK11 (see below for synonymy)
                |    |  i. s.: ‘Daphnella’ aculeola Hedley 1915 [=Tasmadaphne aculeola]P66
                |    |         P. aguayoi (Carcelles 1953)S11
                |    |         P. borbonica Melvill 1923M23
                |    |         *Fusidaphne’ bullata Laseron 1954P66
                |    |         P. eulimines Melv. 1904M23
                |    |         P. gibbera Jeffreys in Bouchet & Waren 1980BW80
                |    |         P. megalembryon (Dautzenberg & Fischer 1896)BW80 (see below for synonymy)
                |    |         P. obesa Bouchet & Waren 1980BW80
                |    |         *Tasmadaphne’ spicula Laseron 1954P66
                |    |         P. thalassica Dall 1919O27
                |    |--P. (Pleurotomella)BK11
                |    |--P. (Anomalotomella Powell 1966)P66
                |    |    |--P. (*A.) anomalapex Powell 1951P66
                |    |    |--P. (A.) annulata Thiele 1912P66
                |    |    |--P. (A.) enderbyensis Powell 1958P66
                |    |    |--P. (A.) frigida Thiele 1912P66
                |    |    `--P. (A.) ohlini (Strebel 1905) [=Thesbia ohlini]P66
                |    |--P. (Cyrtoides Nordsieck 1968)BK11
                |    `--P. (Systenope Cossmann 1889)P66
                |         |--P. (*S.) polycolpa (Cossmann 1889) [=Raphitoma (*S.) polycolpa]P66
                |         |--P. (S.) goniocolpa (Cossmann 1889) [=Raphitoma goniocolpa]P66
                |         |--P. (S.) guepellensis (Cossmann 1889) [=Raphitoma guepellensis]P66
                |         `--P. (S.) linophora (Cossmann 1889) [=Raphitoma linophora]P66
                   `--+--+--Teretiopsis Kanto & Sysoev 1989PK11, BK11
                      |  |    `--T. hyalina Sysoev & Bouchet 2001PK11
                      |  `--+--GymnobelaPK11
                      |     `--Thatcheria Angas 1877PK11, BK11 [incl. Cochlioconus Yokoyama 1928BK11]
                      |          |--T. (Thatcheria)P66
                      |          |    |--*T. (T.) mirabilis Angas 1877P66
                      |          |    |--T. (T.) carinata (Martin 1933) [=Cryptoconus carinatus]P66
                      |          |    |--T. (T.) gradata (Yokoyama 1928) [=*Cochlioconus gradatus]P66
                      |          |    |--T. (T.) liratula (Powell 1942) [=Waitara liratula]P66
                      |          |    |--T. (T.) pagodula (Powell 1942) [=Waitara pagodula]P66
                      |          |    `--T. (T.) vitiensis Charig 1963P66
                      |          `--T. (Waitara Marwick 1931)P66
                      |               `--T. (*W.) waitaraensis (Marwick 1926) [=Turricula waitaraensis]P66
                      `--Daphnella Hinds 1844PK11, BK11 (see below for synonymy)
                           |  i. s.: D. cassandra Hedley 1904H09
                           |         D. cestrum Hedley 1922H22
                           |         D. cheverti Hedley 1922H22
                           |         D. cladara Sysoev & Bouchet 2001BK11
                           |         D. clathrata Gabb 1865O27
                           |         D. crebriplicataH22
                           |         D. cymatodesH22
                           |         D. diluta Sowerby 1896H22
                           |         D. effusa Carpenter 1864C64
                           |         D. electraD19
                           |         D. granata Hedley 1922H22
                           |         D. hayesi Usticke 1959BC01
                           |         D. interrupta Pease 1860K65
                           |         D. maculosa Pease 1860K65
                           |         D. mitrellaformis (Nomura 1940)BK11
                           |         ‘Clathurella’ papuensis Tapparone Canefri 1878H22
                           |         D. pluricarinata (Reeve 1845) [=Pleurotoma pluricarinata]H09
                           |         D. pulvisculus Chino 2006BK11
                           |         D. sigmastoma Hedley 1922H22
                           |         D. stegeriBC01
                           |           |--D. s. stegeriBC01
                           |           `--D. s. monocincta Usticke 1969BC01
                           |         D. stiphra Verco 1909H22
                           |         D. terina Melvill & Standen 1896H22
                           |         D. vitrea (Reeve 1845)P66
                           |--D. (Daphnella)BK11
                           `--D. (Hemidaphne Hedley 1918)BK11
                                |--D. (*H.) souverbiei (Smith 1882) [=Pleurotoma (Daphnella) souverbiei]P66
                                |--D. (H.) axis (Reeve 1846)P66 [=Pleurotoma axisP66, Clathurella axisH22]
                                |--D. (H.) cyclophora (Deshayes 1863)P66 (see below for synonymy)
                                |--D. (H.) lirata (Reeve 1845) [=Pleurotoma lirata]P66
                                |--D. (H.) reeveana (Deshayes 1863)P66 (see below for synonymy)
                                |--D. (H.) rissoides (Reeve 1843) (see below for synonymy)H22
                                |--D. (H.) subula (Reeve 1845) [=Pleurotoma subula]P66
                                |--D. (H.) subuloides Schepman 1913P66
                                `--D. (H.) supracancellata Schepman 1913P66
Raphitomidae incertae sedis:
  Andonia Harris & Burrows 1891 [Andoniinae]BR05
    `--*A. bonellii (Bellardi & Michelotti 1840) [=Fusus bonellii]BR17
  Austrodaphnella Laseron 1954BK11
    `--*A. clathrata Laseron 1954P66
  Hokianga Laws 1947P66
    `--*H. nodulata Laws 1947P66
  Favriella Hornung 1920BK11
    `--*F. weberi (Hornung 1920) [=Daphnella (*Favriella) weberi]P66
  Microgenia Laseron 1954BK11
    `--*M. edwini (Brazier 1894)P66 [=Pleurotoma (Clathurella) edwiniH22, P66, Nepotilla edwiniH22, Taranis edwiniH22]
  Isodaphne Laseron 1954BK11
    `--*I. garrardi Laseron 1954P66
  Fenestrodaphne Powell 1944P66
    `--*F. pulchra Powell 1944P66
  Magnella Dittmer 1960BK11
    `--*M. andersoni Dittmer 1960P66
  Phandella Casey 1903P66
    `--*P. nepionica Casey 1903P66
  Syngenochilus Powell 1944P66
    `--*S. radiapex Powell 1944P66
  Daphnobela Cossmann 1896P66
    `--*D. juncea (Sowerby 1822) [=Buccinum junceum]P66
  Thesbia Jeffreys 1867BK11
    |--*T. nana (Lovén 1846) [=Tritonium nanum]BW80
    `--T. microtoma Cossmann 1899P66
  Leiosyrinx Bouchet & Sysoev 2001BS01
    |--*L. immedicata Bouchet & Sysoev 2001BS01
    |--L. apheles Bouchet & Sysoev 2001BS01
    |--L. liphaima Bouchet & Sysoev 2001BS01
    `--L. matsukumai Bouchet & Sysoev 2001BS01
  Teretia Norman 1888BK11 [incl. Teres Bucquoy, Dautzenberg & Dollfus 1883 non Harris 1829BW80]
    |--*T. teres (Forbes in Reeve 1844) [=Pleurotoma teres; incl. P. boreale Lovén 1846]BW80
    |--T. anceps (Eichwald 1830)P66 (see below for synonymy)
    |--T. nana (Hornung 1920) [=Daphnella (Teres) nana]P66
    `--T. thaumastopsis (Dautzenberg & Fischer 1896) (see below for synonymy)BW80
  Asperdaphne Hedley 1922BK11 [incl. Scabrella Hedley 1918 non Sacco 1890P66]
    |  i. s.: A. perissa (Hedley 1909) [=Mangilia perissa]H22
    |--A. (Asperdaphne)BK11
    `--A. (Aspertilla Powell 1944)BK11
         |--A. (*A.) legrandi (Beddome 1883)P66 (see below for synonymy)
         `--A. (A.) exsculpta Powell 1944P66
  Typhlosyrinx Thiele 1925BS01
    |--*T. vepallida (Martens 1902) (see below for synonymy)BS01
    |--T. neocaledoniensis Bouchet & Sysoev 2001BS01
    |--T. panamica Bouchet & Sysoev 2001BS01
    |--T. praecipua (Smith 1899) [=Pleurotoma (Surcula) praecipua]BS01
    `--T. supracostata (Schepman 1913) [=Surcula supracostata]BS01
  Stilla Finlay 1926BK11
    |--*S. flexicostata (Suter 1899)P66 [=Mangilia flexicostataP66, Nepotilla flexicostataH22]
    |--S. anomala Powell 1955P66
    |--S. delicatula Powell 1927P66
    |--S. fiordlandica Fleming 1948P66
    `--S. paucicostata Powell 1937P66
  Cryptodaphne Powell 1942BK11
    |--*C. pseudodrillia Powell 1942P66
    |--C. abbreviata (Schepman 1913) [=Pleurotomella abbreviata]P66
    |--C. affinis (Schepman 1913) [=Pleurotomella affinis]P66
    |--C. biconica (Schepman 1913) [=Pleurotomella biconica]P66
    `--C. gradata (Schepman 1913) [=Pleurotomella gradata]P66
  Puha Marwick 1931P66
    |--*P. fulgida Marwick 1931P66
    |--P. hebes (Hutton 1873) [=Pleurotoma hebes]P66
    |--P. japonica Shuto 1961P66
    |--P. pusula Laws 1947P66
    `--P. sinusigera (Powell 1942)P66
  Phymorhynchus Dall 1908BK11
    |--*P. castanea (Dall 1895)BW80 (see below for synonymy)
    |--P. alberti (Dautzenberg & Fischer 1906) [=Pleurotoma alberti]BW80
    |--P. argeta (Dall 1889) [=Pleurotomella argeta]P66
    |--P. chevreuxi (Dautzenberg & Fischer 1897)BW80 (see below for synonymy)
    |--P. major Warén & Bouchet 2001BK11
    |--P. oceanica (Dall 1908) [=Pleurotomella (Phymorhynchus) oceanica]P66
    `--P. sulcifera (Bush 1893) (see below for synonymy)BW80
  Pontiothauma Smith 1895BK11
    |--*P. mirabile Smith 1895P66
    |--P. abyssicola Smith 1895P66
    |--P. ergata Hedley 1916P66
    |--P. minus Smith 1906P66
    `--P. pacei Smith 1906P66
  Zenepos Finlay 1928BK11
    |--*Z. totolirata (Suter 1908)P66 [=Daphnella totolirataP66, Nepotilla totolirataH22]
    |--Z. chariessa (Suter 1908) [=Daphnella chariessa]P66
    |--Z. lacunosa (Hutton 1885) [=Daphnella lacunosa]P66
    |--Z. mimica (Sowerby 1896)P66 (see below for synonymy)
    |--Z. minuta (Tenison-Woods 1877)P66 (see below for synonymy)
    `--Z. miranda (Thiele 1925) [=Taranis (Nepotilla) miranda]P66
  Exomilus Hedley 1918BK11
    |--*E. lutrarius (Hedley 1907) [=Mangelia lutraria]P66
    |--E. anxius (Hedley 1909) [=Mangilia anxia]P66
    |--E. ‘cancellatus’ (Beddome 1882)P66 [=Mangilia cancellata non Adams 1865P66, Drillia cancellataH22]
    |--E. cylindricus Laseron 1954P66
    |--E. dyscritos (Verco 1906) [=Terebra dyscritos, Mangilia dyscritos]H22
    |--E. fenestratus (Tate & May 1900) [=Donovania fenestrata, Trophon fenestratus]H22
    |--E. pentagonalis (Verco 1896) [=Drillia pentagonalis]P66
    `--E. telescopialis (Verco 1896) [=Drillia telescopialis]P66
  Teleochilus Harris 1897 [incl. Litachilus Powell 1944]BK11
    |--*T. gracillimus (Tenison-Woods 1877) [=Daphnella gracillima]P66
    |--T. balcombensis Powell 1944P66
    |--T. comptus Powell 1944P66
    |--T. denseliratus Powell 1944P66
    |--T. duplicatus Powell 1944P66
    `--T. royanus Iredale 1924 [=*Litachilus royanus]P66
  Maoridaphne Powell 1942P66
    |--*M. clifdenica (Laws 1939) [=Daphnella clifdenica]P66
    |--M. haroldi Powell 1942P66
    `--M. kaiparica (Laws 1939) [=Daphnella kaiparica]P66
  Spergo Dall 1895BK11
    |--S. (Spergo)BK11
    |    |--*S. (S.) glandiniformis Dall 1895P66
    |    |--S. (S.) daphnelloides Dall 1895P66
    |    |--S. (S.) fusiformis (Kuroda & Habe in Habe 1962)P66, BK11
    |    `--S. (S.) fusus MacNeil 1960P66
    `--S. (Speoides Kuroda & Habe in Habe 1962)BK11
         `--S. (*S.) yoshidai (Kuroda & Habe 1962)P66, BW80 [=*Speoides yoshidaiBW80]
  Clinura Bellardi 1875BK11
    |--*C. calliope (Brocchi 1914) [=Murex (Pleurotoma) calliope]P66
    |--C. anassa Murphy & Rodda 1960P66
    |--C. bituminata (Beets 1843) [=Surculites bituminata]P66
    |--C. controversa (Bellardi 1847) [=Pleurotoma controversa]P66
    |--C. generosa (Marwick 1931) [=Waitara generosa]P66
    |--C. sabatiorum Bellardi 1877P66
    |--C. sopronensis (Wolf 1870) [=Pleurotoma sopronensis]P66
    |--C. subtrochlearis (Friedberg 1912) [=Surcula (Clinura) subtrochlearis]P66
    `--C. trochlearis (Hörnes 1854) [=Pleurotoma trochlearis]P66
  Thatcheriasyrinx Powell 1969BK11
    `--T. orientis (Melvill 1904)BK11
  Rocroithys Sysoiev & Bouchet 2001BK11
    `--R. niveus Sysoev & Bouchet 2001BK11
  Microdaphne McLean 1971BK11
    `--M. morrisoni Rehder 1980BK11
  Abyssobela Kantor & Sysoev 1986BK11
  Abyssothauma Sysoev 1996BK11
  Acamptodaphne Shuto 1971BK11
  Acanthodaphne Bonfitto & Morassi 2006BK11
  Aliceia Dautzenberg & Fischer 1897BK11
    `--*A. aenigmatica Dautzenberg & Fischer 1897BW80 [=Clavus aenigmaticaP66]
  Austropusilla Laseron 1954BK11
    |  i. s.: A. profundis Laseron 1954P66
    |--*A. (Austropusilla) hilum (Hedley 1908)P66 [=Mangelia hilumP66, Filodrillia hilumL54]
    `--A. (Metaclathurella Shuto 1983)BK11
  Bathybela Kobelt 1905 [incl. Bathypota Nordsieck 1968]BK11
    |--*B. nudator (Locard 1897) [=Thesbia nudator]BW80
    `--B. tenellunum (Locard 1897)BW80 (see below for synonymy)
  Cenodagreutes Smith 1967BK11
  Clathurina Melvill 1917BK11
    `--*C. foraminata (Reeve 1845) [=Pleurotoma foraminata, Kermia foraminata]P66
  Famelica Bouchet & Warén 1980BK11
    |--*F. catharinae (Verrill & Smith 1884) [=Famelica catharinae]BW80
    |--F. bitrudis (Barnard 1963) [=Daphnella bitrudis]BW80
    |--F. mirmidina (Dautzenberg & Fischer 1896) [=Pleurotoma mirmidina]BW80
    |--F. monoceros (Watson 1881) (see below for synonymy)BW80
    |--F. monotropis (Dautzenberg & Fischer 1896) [=Pleurotoma monotropis]BW80
    `--F. scipio (Dall 1889) [=Mangilia scipio]BW80
  Kuroshiodaphne Shuto 1965BK11
    |--*K. fuscobalteata (Smith 1879) [=Daphnella fuscobalteata, Maoridaphne (*Kuroshiodaphne) fuscobalteata]P66
    `--K. subzonata (Smith 1879) [=Daphnella subzonata, Maoridaphne (Kuroshiodaphne) subzonata]P66
  Lusitanops Nordsieck 1968BK11 [incl. Pseudazorita Nordsieck 1977BW80]
    |--*L. lusitanica (Sykes 1906) [=Pleurotomella lusitanica, Pleurotomoides (*Lusitanops) lusitanica]BW80
    |--L. blanchardi (Dautzenberg & Fischer 1896) (see below for synonymy)BW80
    |--L. bullioides (Sykes 1906) [=Pleurotomella bullioides]BW80
    |--L. cingulata Bouchet & Waren 1980BW80
    |--‘Pleurotoma’ dyscritaBW80
    |--L. expansa (Sars 1878) [=Bela expansa]BW80
    |--L. hyaloides (Dautzenberg 1925) [=Pleurotoma (Pleurotomella) hyaloides]BW80
    |--L. macrapex Bouchet & Waren 1980BW80
    `--L. sigmoidea Bouchet & Waren 1980BW80
  Mioawateria Vella 1954BK11
    |--*M. personata (Powell 1942) [=Awateria (*Mioawateria) personata]P66
    |--M. expalliata (Laws 1947) [=Awateria (Mioawateria) expalliata]P66
    |--M. experta (Laws 1947) [=Awateria (Mioawateria) experta]P66
    |--M. karakaensis (Marwick 1931) [=Awateria (Mioawateria) karakaensis]P66
    `--M. pahaoaensis (Vella 1954) [=Awateria (Mioawateria) pahaoaensis]P66
  Neopleurotomoides Shuto 1971BK11
    |--*N. rufoapicata (Schepman 1913) [=Clathurella rufoapicata]BW80
    |--N. callembryon (Dautzenberg & Fischer 1896)BW80 (see below for synonymy)
    `--N. distincta Bouchet & Waren 1980BW80
  Pagodidaphne Shuto 1983BK11
  Paramontana Laseron 1954BK11
    |--*P. modesta (Angas 1877)P66 [=Clathurella modestaP66, Pseudodaphnella modestaL54]
    |--P. fusca Laseron 1954P66
    |--P. mayana (Hedley 1922) [=Pseudodaphnella mayana]P66
    |--P. oligoina (Hedley 1922) [=Pseudodaphnella oligoina]P66
    |--P. punicea (Hedley 1922) [=Pseudodaphnella punicea]P66
    `--P. rufozonata (Angas 1877)S11 (see below for synonymy)
  Taranidaphne Morassi & Bonfitto 2001BK11
  Thatcherina Verez-Peláez 1998BK11
  Truncadaphne McLean 1971BK11
  Tuskaroria Sysoev 1988BK11
  Vepridaphne Shuto 1983BK11
  Diaugasma Melvill 1917RO-SF98
    |--*D. epicharta (Melvill & Standen 1903) [=Daphnella (*Diaugasma) epicharta]P66
    |--D. marchadi (Knudsen 1956) [=Philbertia marchadi]RO-SF98
    `--D. olyra (Reeve 1845) [=Pleurotoma olyra, Daphnella (Diaugasma) olyra]P66

Asperdaphne (*Aspertilia) legrandi (Beddome 1883)P66 [=Drillia legrandiP66, Clathurella legrandiH22, Daphnella legrandiH22]

Bathybela tenellunum (Locard 1897)BW80 [=Pleurotoma tenellunumBW80, *Bathypota tenellula (l. c.)BW80, Pleurotomella tenellunumP66; incl. Pleurotomella costlowi Petuch 1974BW80]

Daphnella Hinds 1844PK11, BK11 [incl. Eudaphne Bartsch 1931 non Reuss 1922P66, Eudaphnella Bartsch 1933BK11]

Daphnella (Hemidaphne) cyclophora (Deshayes 1863)P66 [=Pleurotoma cyclophoraP66, Clathurella cyclophoraH22, Mangelia cyclophoraH22]

Daphnella (Hemidaphne) reeveana (Deshayes 1863)P66 [=Pleurotoma reeveanaP66; incl. Clathurella tumida Pease 1867H22]

Daphnella (Hemidaphne) rissoides (Reeve 1843) [=Pleurotoma rissoides, Clathurella rissoides; incl. Pleurotoma dentatum Souverbie 1869 non P. dentata Lamarck 1804, Daphnella dentata, Pusionella testabilis Jousseaume 1896]H22

Famelica monoceros (Watson 1881) [=Pleurotoma (Thesbia) monoceros, Clathurella (Daphnella) monoceros; incl. Mangilia ischna Dall 1927]BW80

Lusitanops blanchardi (Dautzenberg & Fischer 1896) [=Pleurotoma blanchardi, Azorilla blanchardi, *Pseudazorita blanchardi]BW80

Neopleurotomoides callembryon (Dautzenberg & Fischer 1896)BW80 [=Pleurotoma callembryonBW80, Pleurotomella callembryonP66, Pleurotomoides callembryonBW80]

Paramontana rufozonata (Angas 1877)S11 [=Clathurella rufozonataP66, Pseudodaphnella rufozonataL54; incl. Clathurella nodorete May 1915H22, P. rufozonata var. nodoreteH22, Mangelia trachys Tenison-Woods 1878H22, P. rufozonata var. trachysH22]

*Phymorhynchus castanea (Dall 1895)BW80 [=Pleurotomella castaneaBW80; incl. Pl. cingulata Dall 1889BW80, Phymorhynchus cingulataP66, Pl. (Phymorhynchus) clarinda Dall 1908P66, BW80, Ph. speciosus Olsson 1971BW80]

Phymorhynchus chevreuxi (Dautzenberg & Fischer 1897)BW80 [=Pleurotoma chevreuxiBW80, Azorita chevreuxiBW80, Pleurotomella chevreuxiP66]

Phymorhynchus sulcifera (Bush 1893) [=Pleurotomella sulcifera; incl. Pleurotoma adelpha Dautzenberg & Fischer 1896, Pleurotoma fischeri var. costulata Locard 1897 non P. costulatum Cantraine 1835, Pleurotoma decoloratum Locard 1897, Pleurotoma fischeri Locard 1897, Pleurotoma imum Locard 1897, Pleurotoma milneedwardsi Locard 1897]BW80

Pleurotomella Verrill 1873PK11, BK11 [incl. Azorilla Nordsieck 1968BK11, Fusidaphne Laseron 1954BK11, Tasmadaphne Laseron 1954BK11]

Pleurotomella megalembryon (Dautzenberg & Fischer 1896)BW80 [=Pleurotoma megalembryonBW80, *Azorilla megalembryonBW80, Teretia megalembryonP66; incl. Mangilia percompacta Dall 1927BW80]

Teretia anceps (Eichwald 1830)P66 [=Pleurotoma ancepsP66, P. (*Teres) ancepsBW80; incl. P. fusiforme Requien 1848P66, Fusus laviae Calcara 1845P66, P. minutum Aradas 1847P66, Clathurella anceps soluta Marshall 1912P66]

Teretia thaumastopsis (Dautzenberg & Fischer 1896) [=Pleurotoma thaumastopsis; incl. Pleutomella (Eucyclotoma) aperta Dall 1927]BW80

Tritonoturris elegans (Pease 1860) P66 [=Clathurella elegans P66, Defrancia elegans non Möller 1842 (not preoc. if in dif. gen.)]P66

Tritonoturris menecharmes (Melvill 1923)P66 [=Veprecula menecharmesP66; incl. V. menecharmes var. albescens Melvill 1923M23]

*Typhlosyrinx vepallida (Martens 1902) [=Pleurotoma (Leucosyrinx) vepallida, Pleurotoma (Surcula) vepallida, Pleurotomella (Typhlosyrinx) vepallida]BS01

Zenepos mimica (Sowerby 1896)P66 [=Daphnella mimicaP66, Nepotilla mimicaH22; incl. N. mimica var. marmorata Verco 1909H22]

Zenepos minuta (Tenison-Woods 1877)P66 [=Drillia minutaP66, Daphnella minutaH22; incl. Da. mimica var. fusca Sowerby 1896H22]

*Type species of generic name indicated


[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BS01] Bouchet, P., & A. Sysoev. 2001. Typhlosyrinx-like tropical deep-water turriform gastropods (Mollusca, Gastropoda, Conoidea). Journal of Natural History 35: 1693–1715.

[BW80] Bouchet, P., & A. Waren. 1980. Revision of the north-east Atlantic bathyal and abyssal Turridae (Mollusca, Gastropoda). Journal of Molluscan Studies Supplement 8: 1–119.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[D19] Dall, W. H. 1919. Descriptions of new species of mollusks of the family Turritidae from the west coast of America and adjacent regions. Proceedings of the United States National Museum 56: 1–86, pls 1–24.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H22] Hedley, C. 1922. A revision of the Australian Turridae. Records of the Australian Museum 13 (6): 213–359, pls 42–56.

[K65] Kay, E. A. 1965. Marine molluscs in the Cuming collection, British Museum (Natural History) described by William Harper Pease. Bulletin of the British Museum (Natural History): Zoology Supplement 1: 1–96, 14 pls.

[L54] Laseron, C. F. 1954. Revision of the New South Wales Turridae (Mollusca). Royal Zoological Society of New South Wales: Sydney.

[M23] Melvill, J. C. 1923. Descriptions of twenty-one species of Turridae (Pleurotomidae) from various localities in the collection of Mr. E. R. Sykes. Proceedings of the Malacological Society of London 15 (4): 162–171.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[PK11] Puillandre, N., Y. I. Kantor, A. Sysoev, A. Couloux, C. Meyer, T. Rawlings, J. A. Todd & P. Bouchet. 2011. The dragon tamed? A molecular phylogeny of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 259–272.

[RO-SF98] Rolán, E., J. Otero-Schmitt & F. Fernandes. 1998. The family Turridae s.l. (Mollusca, Gastropoda) in Angola (West Africa), 1. Subfamily Daphnellinae. Iberus 16 (1): 95–118.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.


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