Phyllograptus angustifolius elongatus, from here.

Belongs within: Eugraptoloida.
Contains: Isograptus, Bireclinata, Tetragraptinae .

The Reclinata are a clade of graptoloids defined by Maletz et al. (2009) as the smallest clade including Tetragraptus serra and Exigraptus uniformis. Synapomorphies of the clade include the presence of a rutellum on the ventral side of the sicular aperture, serially homologous ventral projections on the proximal thecae (thecal rutellae), and a longer sicula than in outgroup taxa (Maletz et al. 2009).

|--Pan-Bireclinata [Glossograptacea, Glossograptina, Isograptidae]MCM09
| | i. s.: Cardiograptus Harris & Keble in Harris 1916M14
| | |--*C. morsus Harris & Keble 1916B70
| | `--C. crawfordiC73
| | Procardiograptus Xiao, Xia & Wang 1985M14
| | Proncograptus Xiao, Xia & Wang 1985M14
| | Xiushuigraptus Yu & Fang 1983M14
| |--IsograptusMCM09
| `--+--BireclinataMCM09
| `--Parisograptus Chen & Zhang 1996M14
| `--P. caduceus (Salter 1853)M10
`--Tetragraptidae [Pan-Tetragrapta, Tetragrapta]M14
| i. s.: Corymbograptus Obut & Sobolevskaya 1964MCM09, B70
| |--*C. v-fractus (Salter 1863) [=Didymograptus v-fractus]B70
| `--C. retroflexusW04
| Pseudotrigonograptus Mu & Lee 1958MCM09, B70 (see below for synonymy)
| |--*P. uniformis Mu & Lee 1958 [=Tristichograptus uniformis]B70
| |--P. ensiformis (Hall 1859)R93, B70 [=Graptolithus ensiformisB70, *Tristichograptus ensiformisB70]
| `--P. minor Cooper & Fortey 1982R93
| Paratetragraptus Obut 1957M14
| `--*P. approximatus (Nicholson 1873) [=Tetragraptus approximatus]B70
`--+--Pendeograptus Bouček & Přibyl 1951MCM09, B70
| |--*P. pendens (Elles 1898) [=Tetragraptus pendens]B70
| `--P. fruticosus (Hall 1858)MCM09 [=Tetragraptus fruticosusFC86]
`--Phyllograptus Hall 1858MCM09, B70
|--*P. typus Hall 1865B70, M10
|--P. angustifoliusB70
| |--P. a. angustifoliusJ79
| `--P. a. elongatusJ79
|--P. anna Hall 1858R93
|--P. densusJ79
|--P. ilicifolius Hall 1858R93
`--P. loringi White 1874W77

Pseudotrigonograptus Mu & Lee 1958MCM09, B70 [incl. Tristichograptus Jackson & Bulman 1970R93, B70; Pseudotrigonograptidae, Pseudotrigonograptinae]

*Type species of generic name indicated


[B70] Bulman, O. M. B. 1970. Graptolithina with sections on Enteropneusta and Pterobranchia. In: Teichert, C. (ed.) Treatise on Invertebrate Paleontology pt V 2nd ed. pp. V1–V149. The Geological Society of America, Inc.: Boulder (Colorado), and the University of Kansas: Lawrence (Kansas).

[C73] Cooper, R. A. 1973. Taxonomy and evolution of Isograptus Moberg in Australasia. Palaeontology 16: 45–115.

[FC86] Fortey, R. A., & R. A. Cooper. 1986. A phylogenetic classification of the graptoloids. Palaeontology 29: 631–654.

[J79] Jaanusson, V. 1979. Ordovician. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A136–A166. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[M10] Maletz, J. 2010. Xiphograptus and the evolution of virgella-bearing graptoloids. Palaeontology 53 (2): 415–439.

[M14] Maletz, J. 2014. The classification of the Pterobranchia (Cephalodiscida and Graptolithina). Bulletin of Geosciences 89 (3): 477–540.

[MCM09] Maletz, J., J. Carlucci & C. E. Mitchell. 2009. Graptoloid cladistics, taxonomy and phylogeny. Bulletin of Geosciences 84 (1): 7–19.

[R93] Rickards, R. B. 1993. Graptolithina. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 537–542. Chapman & Hall: London.

[W04] Webby, B. D. 2004. Introduction. In: Webby, B. D., F. Paris, M. L. Droser & I. G. Percival (eds) The Great Ordovician Biodiversification Event pp. 1–37. Columbia University Press.

[W77] White, C. A. 1877. Report upon the invertebrate fossils collected in portions of Nevada, Utah, Colorado, New Mexico, and Arizona, by parties of the expeditions of 1871, 1872, 1873, and 1874. U.S. Geographical Surveys West of the One Hundredth Meridian 4 (1): 1–219, pls 1–21.

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