Belongs within: Decapoda.
Contains: Astacidea, Thalassinida, Achelata, Anomala, Brachyura.
The Reptantia are the clade of crustaceans containing decapods other than the shrimps, e.g. crabs, lobsters and crayfish. As the name of the group indicates (‘walkers’), reptants are characterised by more robust legs allowing them to walk rather than swim across the surface of the substrate. The basal Polychelidae retain some features found in natant decapods, such as an elongate, triangular telson and a flexible single articulation between the carpus and propodus of the first pereiopod. The Jurassic to Cretaceous Eryonidae may be early relatives of the Polychelidae. In the Eureptantia, the telson is short and rounded and the first pereiopod bears a stable double hinge between carpus and propodus; Dixon et al. (2003) suggested that the common ancestor of eureptants may have adopted a burrowing habit.
Within the Eureptantia, the Astacura are indicated as the possible sister-group to the remaining taxa (the Sterropoda) by the latter grouping having the first three segmentes of each pereiopod fused, and the third pereiopod lacking a chela (Dixon et al. 2003).
Synapomorphies (from Dixon et al. 2003): Pereiopods articulating in a medio-lateral manner (enabling walking); first two segments of PI fused together. Carapace is calcified and flattened, telson lacking spines. Mandible with long articulation along anterior edge. Third maxilliped shortened relative to other thoracopods; first pereiopod enlarged.
<==Reptantia [Palinura, Palinuridea] |--Polychelida [Eryonoidea]DAS03 | |--EryonidaeBWW93 | | |--Proeryon hargmanni (Von Meyer 1835)BWW93 | | `--Eryon neocomiensis Woodward 1881BWW93 | `--PolychelidaeDAS03 | |--Cardus crucifer (Thomson 1873) [incl. Eryoneicus inermis Bernard 1953]B06 | |--Willemoesiocaris ovalis Van Straelen 1925BWW93 | |--PentachelesB06 | | |--P. laevis Bate 1878 [incl. Polycheles granulatus Faxon 1893]B06 | | `--P. validus Milne Edwards 1880B06 | `--Polycheles [incl. Eryoneicus Bate 1882 (nom. rej.)]B06 | |--P. amemiyai Yokoya 1933B06 | |--P. auriculatus (Bate 1878)MG-H11 | |--P. coccifer Galil 2000MG-H11 | |--P. enthrix (Bate 1878)B06 | |--P. galil Ahyong & Brown 2002DAS03 | |--‘Eryoneicus’ jesperseni Bernard 1953B06 | |--P. nanus (Smith 1884) [=Pentacheles nanus, Stereomastis nana]B06 | |--P. perarmatus Holthuis 1952 [=P. typhlops perarmata]B06 | |--P. sculptus Smith 1880 [=Pentacheles sculptus, Stereomastis sculpta]B06 | |--P. suhmi (Bate 1878)MG-H11 | |--P. talismani Bouvier 1917 [=Polyceles sculptus var. talismani]B06 | `--P. typhlops Heller 1862B06 `--EureptantiaDAS03 |--AstacideaDAS03 `--Sterropoda [Anomura]DAS03 | i. s.: ParalithodesPH03 | |--P. camtschaticus (Tilesius 1815)G10 | `--P. platypus (Brandt 1850)PH03 | AnapagurusKK03 | |--A. bicorniger M-Edwards & Bouvier 1892KK03 | |--A. breviacuelatus Fenizia 1937KK03 | |--A. chiroacanthus (Liljeborg 1856)KK03 | |--A. hyndmanniPP64 | |--A. laevis (Bell 1846)KK03 | |--A. longispina Milne-Edwards & Bouvier 1892KK03 | `--A. petiti Dechancé & Forest 1962KK03 | Australeremus cookiPG98 |--ThalassinidaDAS03 `--EurysternaliaDAS03 |--AchelataDAS03 `--Meiura [Palinurini]DAS03 | i. s.: Mycteris longicarpusH15 | Holoecius cordiformisH15 | Domicea acanthophoraW87 | Lopholithodes mandtiiSBM11 |--AnomalaDAS03 `--BrachyuraDAS03 Reptantia incertae sedis: AcanthacarisDAS03 Chaeraps preissii [incl. C. preissii var. angustus]N24 NotomithraxHS01 |--N. minorHS01 `--N. peroniiHS01 FrediusMT08 |--F. fittkaui (Bott 1967)MT08 `--F. platyacanthus Rodríguez & Pereira 1992MT08 Neosarmatium trispinosumW08 Fabia chilensisFP66 Gourretia denticulata (Lutze 1837)KK03 Pinnaxia transversalisCS77 PotambusCS77 |--P. leptodactylusCS77 `--P. pachypusCS77 Chionocetes opilio (Fabricius 1798)G10 Pugettia quadridens (de Haan 1839)G10 Palaeopalaemon Whitefield 1880S77 [PalaeopalaemonidaeBWW93] |--P. iowensisS74 `--P. newberryi Whitfield 1880BWW93 Cancrinos [Cancrinidae]BWW93 `--C. claviger Münster 1839BWW93 ColeiidaeBWW93 |--Coleia antiquaBWW93 `--HellerocarisBWW93 Tetrachela [Tetrachelidae]BWW93 `--T. raiblana (Bronn 1858)BWW93
*Type species of generic name indicated
References
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