Trident leaf-nosed bats Asellia tridens, photographed by Drew Gardner.

Belongs within: Chiroptera.
Contains: Rhinolophus, Hipposideros.

The Rhinolophidae, Old World leaf-nosed bats, get their vernacular name from the well-developed leaf-like structure at the end of their muzzle.

<==Rhinolophoidea [Yinochiroptera]
    |--+--Rhinopoma Geoffroy 1813M07 (see below for synonymy)
    |  |    |  i. s.: R. cystopsM07
    |  |    |--+--*R. microphyllumM07, FS15 [=Vespertilio microphyllumM07]
    |  |    |  `--R. muscatellumFS15
    |  |    `--+--R. hardwickei Gray 1831FS15, BS72
    |  |       `--R. macinnesiFS15
    |  `--+--CraseonycterisLA02 [CraseonycteridaeMJ11]
    |     |    `--C. thonglongyaiLA02
    |     `--Megadermatidae [Megadermata, Megadermatini, Megadermidae, Megadermina, Megaderminae]MJ11
    |          |  i. s.: Afropterus gigas Lavocat 1961B78
    |          |         Necromantis Weithofer 1887SM93
    |          |           `--N. adichaster Weithofer 1887LA02
    |          |         MiomegadermaKJ70
    |          |         SaharadermaMJ11
    |          |--+--Cardioderma Peters 1873FS15, M07
    |          |  |    `--*C. cor [=Megaderma (*Cardioderma) cor]M07
    |          |  `--Macroderma Miller 1907FS15, HDA88
    |          |       |  i. s.: M. malugara Hand 1996LA02
    |          |       |--M. godthelpi Hand 1985HDA88, LA02
    |          |       `--+--*M. gigas (Dobson 1880)LA02, HDA88, LA02 [=Megaderma gigasM07]
    |          |          |    |--M. g. gigasR64
    |          |          |    `--M. g. saturata Douglas 1962R64
    |          |          `--M. koppa Hand, Dawson & Augee 1988HDA88
    |          `--+--Lavia Gray 1838FS15, M07 [=Livia Agassiz 1846 non Latreille 1802M07]
    |             |    |--*L. frons [=Megaderma (*Lavia) frons]M07
    |             |    `--L. rexM07
    |             `--Megaderma Geoffroy 1810FS15, LA02 (see below for synonymy)
    |                  |--*M. spasma (Linnaeus 1758)LA02 [=Vespertilio spasmaM07, M. (*Spasma) spasmaG66]
    |                  |--M. carimataeM07
    |                  |--M. lyraST01 [=*Lyroderma lyraM07; incl. *Eucheira schistaceaM07]
    |                  |--M. philippinensis Waterhouse 1843W43
    |                  |--M. richardsi Hand 1995LA02
    |                  `--M. trifoliumM07
    `--+--Rhinolophidae [Rhinolophi, Rhinolophineae, Rhinolophinae, Rhinolophini]MJ11
       |    |--RhinolophusKJ70
       |    |--PaleonycterisKJ70
       |    `--RhinomegalophusKJ70
       `--Hipposideridae [Hipposiderinae, Phyllorhinae, Phyllorhininae]HG99
            |  i. s.: Miophyllorhina Hand 1997LA02
            |           `--*M. riversleighensis Hand 1997LA02
            |         Riversleigha Hand 1998LA02
            |           `--*R. williamsi Hand 1998LA02
            |         Xenorhinos Hand 1998LA02
            |           `--*X. halli Hand 1998LA02
            |         Brachipposideros Sigé 1968LA02
            |           |--*B. collongensis (Depéret 1892) [=Hipposideros (*Brachipposideros) collongensis]LA02
            |           |--B. aguilari Legendre 1982LA02
            |           |--B. branssatensis (Hugueney 1965)LA02
            |           |--B. dechaseauxi Sigé 1968LA02
            |           |--B. khengkao Mein & Ginsburg 1997LA02
            |           |--B. nooraleebus Sigé, Hand & Archer 1982LA02
            |           |--B. omani Sigé et al. 1994LA02
            |           `--B. watsoni Hand 1997LA02
            |         PalaeophyllophoriniKJ70
            |           |--PalaeophyllophoraKJ70
            |           `--ParaphyllophoraKJ70
            |         Pseudorhinolophus bouziguensisHG99
            |--Asellia Gray 1838FS15, M07
            |    |--*A. tridens [=Rhinolophus tridens, Hipposideros (*Asellia) tridens, Phyllorhina (*A.) tridens]M07
            |    |--A. mariaetheresae Mein 1958HG99
            |    |--A. murraianaM07
            |    |--A. patriziiIT07
            |    |--A. tricuspidataG66
            |    `--A. vetus Lavocat 1961B78
                 `--+--+----Cloeotis Thomas 1901FS15, M07
                    |  |    `--*C. percivaliM07
                    |  `--+--Rhinonicteris Gray 1847FS15, LA02 [=Rhinonycteris Gray 1866KJ70, M07; Rhinonycterina]
                    |     |    |--*R. aurantius (Gray 1845)LA02 [=Rhinolophus aurantiusM07, *Rhinonycteris aurantiusG66]
                    |     |    `--R. tedfordi Hand 1997LA02
                    |     `--Triaenops Dobson 1871FS15, M07
                    |          |--+--*T. persicus Dobson 1871M07, FS15, K92 [incl. T. aferK92]
                    |          |  `--T. rufusFS15
                    |          `--+--T. auritusFS15
                    |             `--T. furculusFS15
                       |    |--A. stoliczkanusIT07
                       |    `--A. tricuspidatusIT07
                       |         |--A. t. tricuspidatusM72
                       |         `--A. t. novehebridensisM72
                            |--Paracoelops megalotisKJ70, IT07
                            `--Coelops Blyth 1849FS15, B88
                                 |--C. hirsutusFS15
                                 `--+--*C. frithiiM07, FS15 [incl. C. bernsteiniM07]
                                    `--C. robinsoniFS15

Megaderma Geoffroy 1810FS15, LA02 [=Spasma Gray 1866M07; incl. Eucheira Hodgson 1847 non Westwood 1836M07, Lyroderma Peters 1872M07]

Rhinopoma Geoffroy 1813M07 [=Rhinopomus Gervais 1854M07, Rhynopoma Bowdich 1821M07; Rhinopomata, RhinopomatidaeMJ11, Rhinopomatini, Rhinopomidae, Rhinopomina]

*Type species of generic name indicated


[BS72] Bhat, H. R., & M. A. Sreenivasan. 1972. Occurrence of Rhinopoma hardwickei Gray, 1831, the lesser rat-tailed bat in a humid area of coastal Mysore. Journal of the Bombay Natural History Society 69 (1): 216.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[B78] Butler, P. M. 1978. Insectivora and Chiroptera. In: Maglio, V. J., & H. B. S. Cooke (eds) Evolution of African Mammals pp. 56–68. Harvard University Press: Cambridge (Massachusetts).

[FS15] Faurby, S., & J.-C. Svenning. 2015. A species-level phylogeny of all extant and late Quaternary extinct mammals using a novel heuristic-hierarchical Bayesian approach. Molecular Phylogenetics and Evolution 84: 14–26.

[G66] Gray, J. E. 1866. A revision of the genera of Rhinolophidae, or horseshoe bats. Proceedings of the Zoological Society of London 1866: 81–83.

[HDA88] Hand, S., L. Dawson & M. Augee. 1988. Macroderma koppa, a new Tertiary species of false vampire bat (Microchiroptera: Megadermatidae) from Wellington Caves, New South Wales. Records of the Australian Museum 40: 343–351.

[HG99] Hand, S. J., & H. Godthelp. 1999. First Australian Pliocene species of Hipposideros (Microchiroptera: Hipposideridae). Records of the Western Australian Museum Supplement 57: 299–306.

[IT07] Isaac, N. J. B., S. T. Turvey, B. Collen, C. Waterman & J. E. M. Baillie. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PloS One 2 (3): e296.

[K92] Klompen, J. S. H. 1992. Phylogenetic relationships in the mite family Sarcoptidae (Acari: Astigmata). Miscellaneous Publications, Museum of Zoology, University of Michigan 180: i–vi, 1–154.

[KJ70] Koopman, K. F., & J. K. Jones, Jr. 1970. Classification of bats. In: Slaughter, B. H., & D. W. Walton (eds) About Bats pp. 22–28. Southern Methodist University Press: Dallas.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[M72] McKean, J. L. 1972. Notes on some collections of bats (order Chiroptera) from Papua-New Guinea and Bougainville Island. CSIRO Division of Wildlife Research Technical Paper 26: 1–35.

[MJ11] Meredith, R. W., J. E. Janečka, J. Gatesy, O. A. Ryder, C. A. Fisher, E. C. Teeling, A. Goodbla, E. Eizirik, T. L. L. Simão, T. Stadler, D. L. Rabosky, R. L. Honeycutt, J. J. Flynn, C. M. Ingram, C. Steiner, T. L. Williams, T. J. Robinson, A. Burk-Herrick, M. Westerman, N. A. Ayoub, M. S. Springer & W. J. Murphy. 2011. Impacts of the Cretaceous terrestrial revolution and KPg extinction on mammal diversification. Science 334: 521–524.

[M07] Miller, G. S., Jr. 1907. The families and genera of bats. Smithsonian Institution, United States National Museum, Bulletin 57: i–xvii, 1–282, pls 1–14.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1–15.

[ST01] Springer, M. S., E. C. Teeling, O. Madsen, M. J. Stanhope & W. W. de Jong. 2001. Integrated fossil and molecular data reconstruct bat echolocation. Proceedings of the National Academy of Sciences of the USA 98 (11): 6241–6246.

[SM93] Stucky, R. K., & M. C. McKenna. 1993. Mammalia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 739–771. Chapman & Hall: London.

[W43] Waterhouse, G. R. 1843. On various species of bats collected by H. Cuming, Esq. in the Philippine Islands. Proceedings of the Zoological Society of London 11: 66–69.

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