Sacoglossa

Cylindrobulla sp., from Sea Slugs of Hawai’i.

Belongs within: Heterobranchia.
Contains: Plakobranchidae.

The Sacoglossa are a group of algal-feeding gastropods characterised by a muscular, suctorial pharynx and uniseriate radula.

<==Sacoglossa [Sacoglossida] BR17
|–Oxynooidea [Conchoidea, Julioidea, Oxynoacea, Oxynoiformes] BR05
|    |–Cylindrobulla Fischer 1857 BR05 (see below for synonymy)
|    |    |–*C. beauii Fischer 1857 BR17
|    |    |–C. fischeri Adams & Angas 1865 H09
|    |    |–C. gigas Mikkelsen 1998 BC01
|    |    |–C. phuketi Jensen 1989 J96
|    |    `–C. pusilla Nevill 1869 H09
|    |–Oxynoidae [Oxynoeidae] BR05
|    |    |–Oxynoe Rafinesque 1814 BR05
|    |    |    `–*O. olivacea Rafinesque 1814 BR17
|    |    |–Icarus Forbes 1844 [Icaridae, Icarina, Icarinae] BR05
|    |    |    `–*I. gravesi Forbes 1844 BR17
|    |    |–Lophocercus Krohn 1847 [=Lephocercus (l. c.); Lephocercina, Lophocercidae, Lophocercinae] BR05
|    |    |    `–*L. sieboldii Krohn 1847 BR17
|    |    `–Lobiger Krohn 1847 [Lobigeridae, Lobigerina] BR05
|    |         `–*L. philippii Krohn 1847 BR17
|    |–Volvatellidae [Arthessidae, Arthessoidea, Volvatellacea, Volvatellina, Volvatelloidea, Volvatelloidei] BR05
|    |    |–Arthessa Evans 1950 BR05
|    |    |    `–*A. cincta (Nevill & Nevill 1869) [=Volvatella cincta] BR17
|    |    |–Volvatella Pease 1860 [Volvatellinae] BR05
|    |    |    |–*V. fragilis Pease 1860 BR17
|    |    |    `–V. pyriformis Pease 1860 H09
|    |    `–Ascobulla Marcus 1972 [Ascobullacea, Ascobullidae] BR05
|    |         |–*A. ulla (Marcus & Marcus 1970) [=Cylindrobulla ulla] BR17
|    |         `–A. fischeri (Adams & Angas 1864) J96
|    `–Juliidae [Juliacea] BR05
|         |–Gougerotia Le Renard 1980 [Gougerotiinae] BR05
|         |    `–*G. orthodonta Le Renard 1980 BR17
|         |–Berthelinia Crosse 1875 (see below for synonymy) BR05
|         |    |–*B. elegans Crosse 1875 BR17
|         |    `–*Tamanovalva’ limax Kawaguti & Baba 1959 BR17
|         `–Juliinae BR05
|              |–Prasina Deshayes 1863 [Prasinidae] BR05
|              |    `–*P. borbonica Deshayes 1863 BR17
|              `–Julia Gould 1862 BR05
|                   `–*J. exquisita Gould 1862 BR17
|–Plakobranchoidea (see below for synonymy) BR05
|    |–Jenseneria Ortea & Moro 2015 [Jenseneriidae] BR17
|    |    `–*J. borgninii (Trinchese 1896) [=Phyllobranchus borgninii] BR17
|    |–Costasiella Pruvot-Fol 1951 BR05 [Costasiellidae BR17]
|    |    `–*C. virescens Pruvot-Fol 1951 BR17
|    |–Hermaeidae [Caliphyllacea, Caliphyllidae, Caliphyllina, Phyllobranchidae, Phyllobranchillidae] BR17
|    |    |–Caliphylla Costa 1867 BR17, BR05
|    |    |    `–*C. mediterranea Costa 1867 BR17
|    |    |–Phyllobranchillus Pruvot-Fol 1933 [=Phyllobranchus Alder & Hancock 1864 non Girard 1851] BR05
|    |    |    `–*P. orientalis (Kelaart 1858) [=Proctonotus orientalis, *Phyllobranchus orientalis] BR17
|    |    |–Polybranchia Pease 1860 (see below for synonymy) BR05
|    |    |    |–*P. pellucida Pease 1860 [=*Lobifera pellucida] BR17
|    |    |    `–P. westralis Jensen 1993 BW09
|    |    `–Hermaea Lovén 1844 [Hermaeinae] BR05
|    |         |–*H. bifida (Montagu 1815) [=Doris bifida] BR17
|    |         |–H. aoteana Powell 1937 P61
|    |         `–H. dendritica P61
|    |–Limapontiidae [Limapontiadae, Stiligeridae, Stiligerinae, Stiligeroidea] BR05
|    |    |–Calliopaea d’Orbigny 1837 APS-P86
|    |    |–Limapontia Johnston 1836 BR05 [=Pontolimax Creplin 1848 BR17; Limapontiinae, Pontolimacidae]
|    |    |    |–*L. nigra Johnston 1836 BR17
|    |    |    `–L. depressa B64
|    |    |–Olea Agersborg 1923 [Oleidae] BR05
|    |    |    `–*O. hansineensis Agersborg 1923 BR17
|    |    |–Alderia Allmann 1845 [Alderiidae] BR05
|    |    |    `–*A. modesta (Lovén 1844) [=Stiliger modestus] BR17
|    |    |–Ercolania Trinchese 1872 [Ercolaniinae] BR05
|    |    |    |–*E. siottii Trinchese 1872 BR17
|    |    |    `–E. funerea (Costa 1867) H01
|    |    `–Stiliger Ehrenberg 1828 BR05
|    |         |–*S. ornatus Ehrenberg 1828 BR17
|    |         |–S. bellulus [=Calliopaea bellula] P61
|    |         |–S. felinus (Hutton 1882) P61
|    |         `–S. vossi CG99
|    `–Plakobranchidae BR05
`–Platyhedylidae [Platyhedylacea, Platyhedyloidea] BR17
|–Gascoignella Jensen 1985 [Gascoignellidae] BR05
|    `–*G. aprica Jensen 1985 BR17
`–Platyhedyle Salvini-Plawen 1973 BR05
`–*P. denudata Salvini-Plawen 1973 BR17

Berthelinia Crosse 1875 [incl. Tamanovalva Kawaguti & Baba 1959; Bertheliniidae, Bertheliniinae, Tamanovalvacea, Tamanovalvidae, Tamanovalvoidei] BR05

Cylindrobulla Fischer 1857 BR05 [Cylindrobullacea, Cylindrobullida, Cylindrobullidae BR17, Cylindrobullina, Cylindrobullinae, Cylindrobulloidea]

Plakobranchoidea [Aconchoidea, Ascoglossa, Cirrobranchia, Elysiacea, Elysiina, Elysiomorpha, Limapontioidea, Pellibranchiata, Phyllobranchiacea, Placobranchacea, Placobranchoidea, Plakobranchacea, Stiligerida, Stiligerina] BR05

Polybranchia Pease 1860 [=Lobifera Pease 1860; Lobiferidae, Polybranchidae, Polybranchiidae, Polybranchiinae, Polybranchioidea] BR05

*Type species of generic name indicated

REFERENCES

[APS-P86] Arnaud, P. M., C. Poizat & L. v. Salvini-Plawen. 1986. Marine-interstitial Gastropoda (including one freshwater interstitial species). In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 153–176. E. J. Brill/Dr W. Backhuys: Leiden.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H01] Huys, R. 2001. Splanchnotrophid systematics: a case of polyphyly and taxonomic myopia. Journal of Crustacean Biology 21 (1): 106–156.

[J96] Jensen, K. R. 1996. The Diaphanidae as a possible sister group of the Sacoglossa (Gastropoda, Opisthobranchia). In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 231–247. Oxford University Press: Oxford.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

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