Scaphisoma agaricinum, copyright Siga.

Belongs within: Staphylinidae.

The Scaphidiinae are a group of rove beetles with a convex body form and long elytra covering most of the abdomen. As far as is known, all species are mycophagous (Newton et al. 2001).

Characters (from Newton et al. 2001): Body form broadly oval, compact, very convex and shiny, with long slender legs and long but truncate elytra that leave only 1–2 abdominal segments fully exposed; head small, somewhat retracted and hypognathous, antennae inserted on frons between eyes, neck absent; procoxal fissure very short and narrow to absent, protrochantin always concealed; abdomen more or less conical, with six visible sterna and 0–2 pairs of paratergites per segment; tarsi 5-5-5 .

<==Scaphidiinae [Scaphidiidae, Scaphidites]
| i. s.: Scaphidiopsis aequivocaLC20
|--Brachynopus latusLC20
`--+--Scaphidiini [Cerambyciscaphini, Diateliini]B14
| |--Diatelium wallacei Pascoe 1863B14
| `--Scaphidium Olivier 1790LC20, NT01
| | i. s.: S. alpicola Blackb. 1891M96
| | S. atomariumR26
| | S. basale Laporte de Castelnau 1840LC40
| | S. bicolor Laporte de Castelnau 1840LC40
| | S. consobrinum Laporte de Castelnau 1840LC40
| | S. fasciatum Laporte de Castelnau 1840LC40
| | S. deletumLC20
| | S. immaculatumL02
| | S. mexicanum Laporte de Castelnau 1840LC40
| | S. nigrum Laporte de Castelnau 1840LC40
| | S. pardalis Laporte de Castelnau 1840LC40
| | S. piceumNT01
| | S. punctatum Laporte de Castelnau 1840LC40
| | S. punctipenneLB91
| | S. quadrinotatumLC40
| | S. quadripustulatumB70
| | S. simile Laporte de Castelnau 1840LC40
| | S. unicolor Laporte de Castelnau 1840LC40
| |--S. quadrimaculatumLC20
| `--+--S. celebenseLC20
| `--S. picconiiLC20
`--+--Scaphium Kirby 1837LC20, NT01 [Scaphiini]
| `--S. castanipes Kirby 1837NT01
`--+--Cyparium Erichson 1845LC20, NT01 [Cypariini]
| |--C. ater Casey 1900NT01
| `--C. concolor (Fabricius 1801) [incl. C. flavipes LeConte 1860]NT01
`--Scaphisomatini [Baeoceridiina, Baeocerina, Heteroscaphini, Scaphisomini, Toxidiini]NT01
| i. s.: Caryoscapha Ganglbauer 1899NT01
| `--C. americanum Löbl 1987NT01
| Toxidium LeConte 1860NT01
| |--T. compressum Zimmermann 1869NT01
| `--T. gammaroides LeConte 1860NT01
|--+--Baeocera Erichson 1845LC20, NT01 [incl. Eubaeocera Cornell 1967NT01, Sciatrophes Blackburn 1903NT01]
| `--XotidiumLC20
| `--ScaphicomaLC20
| |--S. hiranoiLC20
| `--+--S. bidentiaLC20
| `--S. quadrifasciataLC20
`--+--Bironium amicaleLC20
`--+--+--+--Sapitia versicolorLC20
| | `--TermitoscaphiumLC20
| `--+--Vituratella termitophilumLC20
| `--Baeoceridium celebenseLC20
`--Scaphisoma Leach 1815LC20, NT01 [incl. Scaphiomicrus Casey 1900NT01]
| i. s.: S. agaricinumC01 [=Scaphidium agaricinumLC40]
| S. assimileC01
| S. castaneumH96
| S. communeNT01
| S. gracileLC20
| S. impunctatumH96
| S. novicum Blackb. 1891M96
|--S. latitarseLC20
`--+--S. napuLC20
`--S. paluLC20

*Type species of generic name indicated


[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[H96] Hanley, R. S. 1996. Immature stages of Scaphisoma castaneum Motschulsky (Coleoptera: Staphylinidae: Scaphidiinae), with observations on natural history, fungal hosts and development. Proceedings of the Entomological Society of Washington 96 (1): 36–43.

[LC40] Laporte de Castelnau, F. 1840. Histoire Naturelle des Insectes Coléoptères vol. 2. P. Duménil: Paris.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LC20] Lü, L., C.-Y. Cai, X. Zhang, A. F. Newton, M. K. Thayer & H.-Z. Zhou. 2020. Linking evolutionary mode to palaeoclimate change reveals rapid radiations of staphylinoid beetles in low-energy conditions. Current Zoology 66 (4): 435–444.

[M96] Masters, G. 1896. Catalogue of the described Coleoptera of Australia. Supplement, part II. Dytiscidae, Gyrinidae, Hydrophyllidae, Staphylinidae, Pselaphidae, Paussidae, Silphidae, Scaphididae, Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydiidae, Cucujidae, Cryptophagidae, Lathridiidae, Mycetophagidae, Dermestidae, Byrrhidae, Parnidae, Heteroceridae. Proceedings of the Linnean Society of New South Wales 21 (Suppl.): 695–754.

[NT01] Newton, A. F., M. K. Thayer, J. S. Ashe & D. S. Chandler. 2001. Staphylinidae Latreille, 1802. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 272–418. CRC Press: Boca Raton.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

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