Scaphites

Reconstruction of female Scaphites preventricosus, from Landman et al. (2017).

Belongs within: Scaphitidae.

Scaphites is a Cretaceous (Upper Albian to Campanian) genus of heteromorph ammonites (Wright et al. 1996).

Characters (from Wright et al. 1996): Compressed to very inflated; spire more or less involute; early whorls in contact; shaft short to moderately long; hook not curved over spire; aperture constricted and commonly collared, in some with long dorsal lappet; ribs on spire normally long and short or branching; ribs on shaft single or branching, commonly at ventrolateral tubercles; umbilical and ventrolateral tubercles may be present on shaft and hook. Suture with L initially bifid but becoming asymmetric in Upper Turonian and later trifid; saddle U/L very wide and divided by one or more auxiliary lobes.

<==Scaphites Parkinson 1811EK02 (see below for synonymy)
    |--*S. equalis Sowerby 1813 [=*Holcoscaphites equalis]WCH96
    |--S. arcadiensisK79
    |--S. binneyiK79
    |--S. carlilensisK79
    |--S. corvensisK79
    |--S. depressusK79
    |--S. eruciformis Etheridge 1902F71
    |--S. ferronensisK79
    |--S. frontierensisK79
    |--S. geinitzi [incl. S. geinitzi var. binodosus Jahn 1891 non S. binodosus Roemer 1841, *Jahnites binodosus]WCH96
    |--S. hippocrepisW69
    |--S. leeiK79
    |--S. mariasensisK79
    |--S. multicostatumK79
    |--S. nicolleti (Morton 1842) [=Hoploscaphites nicolleti]CF06
    |--S. nigricollensisW69
    |--S. obliquusW69
    |--S. planusK79
    |--S. preventricosusTLY03
    |--S. reesideiK79
    |--S. simplexW69
    |--S. subdelicatulus Cobban & Gryc 1961 [=S. subdelicatus (l. c.)]HK81
    |--S. thomasi Pervinquière 1907 [=*Metascaphites thomasi]WCH96
    |--S. ventricosus Meek & Hayden 1863 [=*Anascaphites ventricosus]WCH96
    |--S. warreni Meek & Hayden 1860W77
    `--S. whitfieldiTLY03

Scaphites Parkinson 1811EK02 [=Holcoscaphites Nowak 1911WCH96; incl. Anascaphites Hyatt 1900WCH96, Jahnites Hyatt 1900WCH96, Metascaphites Wiedmann 1962WCH96]

*Type species of generic name indicated

References

[CF06] Crawford, R. S., R. F. Feldmann, D. A. Waugh, B. M. Kelley & J. G. Allen. 2006. Decapod crustaceans from the Maastrichtian Fox Hills formation. Bulletin of the Peabody Museum of Natural History 47 (1–2): 3–28.

[EK02] Engeser, T., & H. Keupp. 2002. Phylogeny of the aptychi-possessing Neoammonoidea (Aptychophora nov., Cephalopoda). Lethaia 35: 79–96.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[HK81] Hancock, J. M., & W. J. Kennedy. 1981. Upper Cretaceous ammonite stratigraphy: some current problems. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 531–553. Academic Press.

[K79] Kauffman, E. G. 1979. Cretaceous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A418–A487. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[TLY03] Tanabe, K., N. H. Landman & Y. Yoshioka. 2003. Intra- and interspecific variation in the early internal shell features of some Cretaceous ammonoids. Journal of Paleontology 77 (5): 876–887.

[W77] White, C. A. 1877. Report upon the invertebrate fossils collected in portions of Nevada, Utah, Colorado, New Mexico, and Arizona, by parties of the expeditions of 1871, 1872, 1873, and 1874. U.S. Geographical Surveys West of the One Hundredth Meridian 4 (1): 1–219, pls 1–21.

[W69] Wiedmann, J. 1969. The heteromorphs and ammonoid extinction. Biological Reviews 44: 563–602.

[WCH96] Wright, C. W., J. H. Calloman & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

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