Big-headed ground beetle Scarites subterraneus, copyright Kaldari.

Belongs within: Carabidae.
Contains: Laccopterum, Scaraphites, Conopterum, Neocarenum, Monocentrum, Euryscaphus, Carenum, Dyschirius, Clivina, Dyschiriodes.

The Scaritinae are a diverse group of ground beetles with body forms commonly modified for digging. The body is pedunculate with a marked constriction between pro- and mesothorax. The fore coxal cavities are closed behind and the mid coxal cavities laterally open, partially closed by the mesepimeron (Lawrence & Britton 1991). The tribe Clivinini includes relatively small forms with a prominent, setiform unguitractor plate on the tarsus whereas members of the Scaritini have a much smaller plate and tend to be much larger in overall size (Ball & Bousquet 2001).

Scaritinae [Scarititae, Scrobifera] B14
    |–Migadopini [Migadopinae] R-JC01
    |    |–Migadops latus R-JC01
    |    `–Stichonotus Sloane 1910 LB91, LM87
    |         |–*S. leai Sloane 1910 LM87
    |         |–S. limbatus Sloane 1915 LM87
    |         `–S. piceus Sloane 1915 LM87
    |–Scaritini B90
    |    |–Euryscaphus LB91
    |    |–Carenum LB91
    |    |–Dyschirius B90
    |    |–Pasimachus Bonelli 1813 BB01
    |    |    |  i. s.: P. elongatus LeConte 1846 BB01
    |    |    |         P. subangulatus (Chaudoir 1862) B14
    |    |    `–P. (Emydopterus Lacordaire 1854) BB01
    |    `–Scarites Fabricius 1775 BB01
    |         |–S. eurytus G89
    |         |–S. gigas C74
    |         |–S. guineensis [incl. S. guineensis var. asphaltinus] G89
    |         |–S. pyracmon B35
    |         |–S. rugosus B35
    |         |–S. senegalensis B55
    |         `–S. (Scarites) subterraneus Fabricius 1775 BB01
    `–Clivinini BB01
         |–Clivina BB01
         |–Ardistomis Putzeys 1846 BB01
         |–Dyschiriodes BB01
         |–Oxydrepanus Putzeys 1866 BB01
         |    `–O. rufus (Putzeys 1846) BB01
         |–Halocoryza Alluaud 1919 BB01
         |    `–H. arenaria (Darlington 1939) BB01
         |–Aspidoglossa Putzeys 1846 BB01
         |    `–A. subangulata (Chaudoir 1843) BB01
         |–Semiardistomis Kult 1950 BB01
         |    |–S. puncticollis (Dejean 1831) BB01
         |    `–S. viridis (Say 1823) BB01
         |–Akephorus LeConte 1852 BB01
         |    |–A. marinus LeConte 1852 BB01
         |    `–A. obesus LeConte 1866 B14
         `–Schizogenius Putzeys 1846 BB01
              |  i. s.: S. freyi Baehr 1983 BB01
              `–S. (Genioschizus Whitehead 1972) BB01
                   `–S. (G.) crenulatus LeConte 1852 BB01

Scaritinae incertae sedis:
  Ochyropus gigas B55
  Taeniolobus B55
    |–T. laevipennis B55
    |–T. lucidus B55
    |    |–T. l. lucidus B55
    |    `–T. l. strigiceps B55
    `–T. vilhenai Basilewsky 1955 B55
  Trilophus congoanus B55
  Syleter Andrewes 1941 [=Psilus Putzeys 1877 nec Panzer 1806 nec Fischer de Waldheim 1813] LM87
    |–*S. paradoxus (Putzeys 1868) [=Ardistomis paradoxa] LM87
    |–S. andrewesi (Basilewsky 1931) [=Dyschirius andrewesi, Psilus andrewesi] B55
    |–S. congoensis B55
    `–S. papua Darlington 1962 LM87
  Pseudoclivina Kult 1947 B55
    `–P. grandis B55
  Rhysocara Sloane 1916 LM87
    `–*R. crassa Sloane 1916 LM87
  Platysphyrus Sloane 1905 LM87
    `–*P. tibialis Sloane 1905 LM87
  Clivinarchus Sloane 1896 LM87
    `–*C. perlongus Sloane 1896 LM87
  Laccopterum LM87
  Philoscaphus Macleay 1871 LM87
    |–*P. tuberculatus (Macleay 1863) [=Carenum tuberculatum] LM87
    |–P. barnardi Macleay 1888 LM87
    |–P. bicostatus Sloane 1905 LM87
    |–P. carinatus (Macleay 1864) [=Carenum carinatum] LM87
    |–P. costalis Macleay 1873 (see below for synonymy) LM87
    `–P. mastersii Macleay 1871 [incl. P. duboulayi Blackburn 1890] LM87
  Scaraphites LM87
  Conopterum LM87
  Neocarenum LM87
  Trichocarenum Blackburn 1892 LM87
    |–*T. elderi Blackburn 1892 LM87
    |–T. castelnaui Sloane 1905 LM87
    `–T. cylindricum Sloane 1897 LM87
  Neoscaphus Sloane 1888 LM87
    `–*N. simplex Sloane 1888 LM87
  Epilectus Blackburn 1888 [=Carenarchus Sloane 1905, Eurygnathus Blackburn 1888 non Wollaston 1854] LM87
    |–*E. fortis (Blackburn 1888) [=*Eurygnathus fortis, *Carenarchus fortis] LM87
    `–E. mastersi (Macleay 1869) [=Neocarenum mastersi] LM87
  Monocentrum LM87
  Steganomma Macleay 1887 LM87
    `–*S. porcatum Macleay 1887 LM87
  Geoscaptus Chaudoir 1855 LM87
    |–*G. laevissimus Chaudoir 1855 (see below for synonymy) LM87
    |–G. cacus (Macleay 1863) (see below for synonymy) LM87
    |–G. crassus Sloane 1895 LM87
    |–G. macleayi Chaudoir 1879 LM87
    `–G. plicatulus (Castelnau 1867) [=Scarites plicatulus] LM87
  Omoglymmius B14
    |–O. americanus (Laporte 1836) B14
    `–O. hamatus B14
  Solenogenys funkei Adis 1981 B14

Geoscaptus cacus (Macleay 1863) [=Scarites cacus; incl. S. approximatus Macleay 1865, S. bipunctatus Castelnau 1867, S. mitchellii Castelnau 1867, S. ruficornis Castelnau 1867, S. subporcatulus Macleay 1865, S. substriatus Castelnau 1867, S. waterhousei Macleay 1865] LM87

*Geoscaptus laevissimus Chaudoir 1855 [incl. Scarites bostocki Castelnau 1867, S. damastes Macleay 1863, S. geryon Macleay 1863, S. jacksoniensis Macleay 1865, S. planiusculus Macleay 1865] LM87

Philoscaphus costalis Macleay 1873 [incl. P. crassus Blackburn 1888, P. lateralis Macleay 1873, P. tepperi Blackburn 1888] LM87

*Type species of generic name indicated


[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C74] Coineau, Y. 1974. Éléments pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). Mémoires du Muséum National d’Histoire Naturelle, nouvelle série, Série A, Zoologie 81: 1–299, 24 pls.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Last updated: 19 December 2020.

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