Scenellidae

Latex mould of Pichynella annulata, from Tortello & Sabattini (2011). Scale bar = 0.2 mm.

Belongs within: Helcionellida.

The Scenellidae are a group of univalved molluscs known from the Early Cambrian, with a simple cap-shaped shell without a peripheral buttress (Parkhaev 2002, as Helcionellidae).

Scenellidae [Helcionellidae]BR05
|--Yangtzeconinae [Yangtzeconiacea, Yangtzeconidae]BR17
| |--Yangtzeconus Yu 1979BR17
| | `--*Y. priscus Yu 1979BR17
| `--Ceratoconus Chen & Zhang 1980 [Ceratoconidae]BR17
| `--*C. striatus Chen & Zhang 1980BR17
`--Scenellinae (see below for synonymy)BR17
|--Palaeacmaea Hall & Whitfield 1872BR17, KY60
| |--*P. typica Hall & Whitfield 1872KY60
| `--P. solarium Lindström 1884K52
|--Helcionella Grabau & Shimer 1909P02
| `--H. subrugosa (d’Orbigny 1850)KC60 (see below for synonymy)
| |--H. s. subrugosaRJ76
| `--H. s. comleyensisRJ76
|--Hampilina Kobayashi 1958BR17, P02 [Hampilininae, Hamplininae]
| `--*H. goniospira Kobayashi 1958KC60
|--Eosoconus Yu 1979 [Eosoconidae]BR17
| `--*E. primarius Yu 1979BR17
|--Kundatoides Aksarina 1995Y08
| `--K. perplexus Aksarina 1995Y08
|--Davidella Zhegallo 1996Y08
| `--*D. tommotica Zhegallo 1996Y08
|--Merismoconcha Yu 1979 [Merismoconchinae]Y08
| `--*M. multisegmentata Yu 1979Y08
|--Actinoconus Yu 1979 [Actinoconidae]BR17
| `--*A. pyriformis Yu 1979BR17
|--Yangtzemerisma Yu 1984Y08
| `--*Y. rarum Yu 1984 [incl. Y. cancellata Yu 1984]Y08
|--Minymerisma Yu 1984Y08
| `--*M. yunnanense Yu 1984Y08
|--Marocella Geyer 1986BR17, P02 [Marocellidae]
| `--*M. mira Geyer 1986BR17
|--Shelbyoceras Ulrich & Foerste 1936 [Shelbyoceratidae, Shelbyoceridae]BR17
| |--*S. robustum Ulrich & Foerste 1936BR17
| |--S. bessemerense Ulrich, Foerste & Miller 1943P68
| |--S. buttsi Ulrich, Foerste & Miller 1943P68
| `--S. unguliforme Ulrich, Foerste & Mller 1943P68
|--Scenella Billings 1872P02 [incl. Parmophorella Matthew 1886K52]
| |--*S. reticulata Billings 1872F62
| |--S. amiiTS11
| |--S. barrandeiTS11
| |--S. jijiapoensis Chen 1984NS93
| |--S. micropora He 1984NS93
| `--S. radiansTS11
`--Calloconus Perner 1903K52
`--*C. humilis (Perner 1903) [=Palaeoscurria (*Calloconus) humilis]KY60

Scenellidae incertae sedis:
Bemella Missarzhevsky in Rozanov et al. 1969 [incl. Charaulachella Vassiljeva 1990]P02
|--B. jacutica Missarzhevsky 1969D94
`--B. septataP08
Ilsanella Missarzhevsky 1981P02
`--I. atdabanicaP08
Tannuella Missarzhevsky in Rozanov et al. 1969P02
`--T. elata Missarzhevskii 1969BE93
Igorella Missarzhevsky in Rozanov et al. 1969P02
|--I. emeiensisP08
|--I. maidipingensisP08
`--I. monstrosaRJ76
Obtusoconus Yu 1979P02
|--O. brevisP08
`--O. honorabilisP08
Emarginoconus Yu 1979P02
`--E. mirusP08
Igorellina Missarzhevsky 1989P02
`--I. monstrosaP08
Pararaconus Runnegar in Bengtson et al. 1990P02
Lenoconus Vassiljeva 1990P02
`--L. sulcatusP08
Asperconella Landing in Landing & Bartowski 1996P02
Pseudopatella Zhegallo in Esakova & Zhegallo 1996 nec Sacc. 1884 (ICBN) nec Speg. 1891 (ICBN)P02
Paucella Vassiljeva 1998P02
Randomia Matthew 1899P02
`--*R. aurorae Matthew 1899F62
Pichynella Rusconi 1954F62
`--*P. annulata Rusconi 1954F62
Totoralia Tortello & Sabattini 2011TS11
`--*T. hornillensis (Rusconi 1956) [=Pichynella hornillensis; incl. P. striata Rusconi 1958]TS11
‘Ginella’ Missarzhevsky in Rozanov et al. 1969 non Ivanova 1960TS11
`--G. savitzkiiRJ76
AbsidaticonusP08
Aequiconus zigzacP08
Anuliconus magnificusP08
CalyptroconusP08
Chabaktiella Missarzhevsky 1981P08, P02
ChuiliellaP08
CodonoconusP08
Daedalia daedalaP08
Fenqiaronia proboscisP08
MiroconulusP08
PseudoscenellaP08
SalanyellaP08
Truncatoconus campylurusP08
TuoraconusP08
Aegitellus placusP08
Auricullina papulosaP08
Cycloholcus Knight 1947RJ76
`--*C. nummus Knight 1947KC60
Pollicina Koken in Holzapfel 1895RJ76, BR17 [Pollicinidae]
|--*P. corniculum (Eichwald 1860) (see below for synonymy)BR17
`--P. cambrica (Moberg 1892) [=Dentalium cambricum]MB-M99
Pseudopollicina Vostokova 1962RJ76
‘Lenaella’ Byalyy 1973 non Korde 1959RJ76
Macroscenella Wilson 1951RJ76
`--*M. superba (Billings 1865) [=Metoptoma superba]KY60
Palaeoscurria Perner 1903RJ76
|--*P. calyptrata Perner 1903KC60
`--P. ordoviciana Horný 1961TTE93
Platypilina Horny 1961RJ76
Pseudomatthevia Shaw 1956RJ76

Helcionella subrugosa (d’Orbigny 1850)KC60 [=Helcion subrugosusP08, Metoptoma rugosa Hall 1847 non Patella rugosa Sowerby 1816KC60, Helcion rugosaKC60, *Helcionella rugosaBR17]

*Pollicina corniculum (Eichwald 1860) [=Cyrtolites corniculum, Cyrtoceras laeve Eichwald 1842 non Sowerby 1839, Cyrtolites laevis]BR17

Scenellinae [Helcionellinae, Merismoconchia, Merismoconchiacea, Merismoconchida, Merismoconchidae, Merismoconchioidea, Palaeacmaeidae, Palaeacmaeinae, Yangtzemerismatinae]BR17

*Type species of generic name indicated

References

[BE93] Benton, M. J., & D. H. Erwin. 1993. Mollusca: Amphineura and ‘Monoplacophora’. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 125–130. Chapman & Hall: London.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[D94] Dzik, J. 1994. Evolution of ‘small shelly fossils’ assemblages of the early Paleozoic. Acta Palaeontologica Polonica 39 (3): 247–313.

[F62] Fisher, D. W. 1962. Small conoidal shells of uncertain affinities. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt W. Miscellanea: Conodonts, Conoidal Shells of Uncertain Affinities, Worms, Trace Fossils and Problematica pp. W98–W143. Geological Society of America, and University of Kansas Press.

[K52] Knight, J. B. 1952. Primitive fossil gastropods and their bearing on gastropod classification. Smithsonian Miscellaneous Collections 117 (13): 1–56, 2 pls.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[KY60] Knight, J. B., & E. L. Yochelson. 1960. Monoplacophora. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I77-I84. Geological Society of America, and University of Kansas Press.

[MB-M99] Malinky, J. M., & V. Berg-Madsen. 1999. A revision of Holm’s Early and early Mid Cambrian hyoliths of Sweden. Palaeontology 42: 25–65.

[NS93] Nudds, J. R., & J. J. Sepkoski Jr. 1993. Coelenterata. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 101–124. Chapman & Hall: London.

[P02] Parkhaev, P. Yu. 2002. Phylogenesis and the system of the Cambrian univalved mollusks. Paleontologicheskii Zhurnal 2002 (1): 27–39 (translated: Paleontological Journal 36 (1): 25–36).

[P08] Parkhaev, P. Y. 2008. The Early Cambrian radiation of Mollusca. In: Ponder, W. F., & D. R. Lindberg (eds) Phylogeny and Evolution of the Mollusca pp. 33–69. University of California Press: Berkeley.

[P68] Purnell, L. R. 1968. Catalog of the type specimens of invertebrate fossils. Part I: Paleozoic Cephalopoda. United States National Museum Bulletin 262: 1–198.

[RJ76] Runnegar, B., & P. A. Jell. 1976. Australian Middle Cambrian molluscs and their bearing on early molluscan evolution. Alcheringa 1 (2): 109–138.

[TS11] Tortello, M. F., & N. M. Sabattini. 2011. Totoralia, a new conical-shaped mollusk from the Middle Cambrian of western Argentina. Geologica Acta 9 (2): 175–185.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[Y08] Yu W. 2008. On the genus Yangtzemerisma and related genera (Mollusca: Merismoconchia). Records of the Western Australian Museum 24 (2): 181–194.

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