Specimens of Schwagerina huecoensis, from here.

Belongs within: Fusulinoida.

The Schwagerininae are a group of fusulinid Foraminifera known from the Upper Carboniferous and Permian with outer test walls comprised of two layers. The test is tightly coiled in most genera, but becomes irregularly and loosely uncoiled in gerontic stages of the Japanese Permian genus Nipponitella (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Shell large, fusiform to irregularly cylindrical, planispiral, involute in most, irregularly uncoiled in some; spirotheca thick, composed of tectum and alveolar keriotheca; septa fluted in end zones of primitive genera, fluted completely across shell and to tops of chambers of more advanced genera; tunnel singular in most forms or multiple in one genus, axial fillings absent to massive; chomata massive to slight.

    |--Alpinoschwagerina Bensh 1972R-CB96
    |    `--*A. turkestanica Bensh 1972R-CB96
    |--Rugososchwagerina Miklukho-Maclay 1956R-CB96
    |    |--*R. yabei (Staff 1909) [=Schwagerina yabei]LT64
    |    `--R. fosteriLT64
    |--Sphaeroschwagerina Miklukho-Maklay 1959LT64
    |    |--S. sphaerica Scherbovich 1949 (see below for synonymy)R-CB96
    |    |--S. fusiformisRR79
    |    |--S. karnica (Shcherbovich in Rauzer-Chernousova & Shcherbovich 1949) [=Schwagerina sphaerica var. karnica]LT64
    |    |--S. moelleriRR79
    |    `--S. vulgarisRR79
    `--Schwagerina Moeller 1877R-CB96 [incl. Codonoschwagerina Viên 1959LT64]
         |--S. moelleri Rauser 1936 [=*S. princeps Moeller 1877 non Borelis princeps Ehrenberg 1842]R-CB96
         |--S. arctica Xia in Ding, Xia et al. 1984DX84
         |--S. bulegensis Xia in Ding, Xia et al. 1984DX84
         |--S. chihsiaensisDX84
         |--S. confertaDX84
         |--S. conicocylindricaDX84
         |--S. crassitectoriaRR79
         |--S. cushmaniCB86
         |--S. douvelleiDX84
         |--S. dugoutensisRR79
         |--S. faxLT64
         |--S. furoniLT64
         |--S. granumavenaeDX84
         |--S. gruperaensisLT64
         |--S. hawkinsiRR79
         |--S. huecoensisC40
         |--S. hupehensisDX84
         |--S. kwangchiensis Chen 1956DX84
         |--S. longaDX84
         |--S. longipertica Chen 1956DX84
         |--S. mandulaensis Xia in Ding, Xia et al. 1984DX84
         |--S. moungthensisG31
         |--S. neimonggolensis Xia in Ding, Xia et al. 1984DX84
         |--S. normalis Xia in Ding, Xia et al. 1984DX84
         |--S. pactirugaDX84
         |--S. princeps (Ehrenberg 1842) [=Borelis princeps]LT64
         |--S. pseudoregularisDX84
         |--S. puerilis Xia in Ding, Xia et al. 1984DX84
         |--S. pulchella Xia in Ding, Xia et al. 1984DX84
         |--S. quasiregularis Sheng 1963DX84
         |--S. robustaG31
         |--S. rutschiLT64
         |--S. setumRR79
         |--S. thuanae (Viên 1959) [=*Codonoschwagerina thuanae]LT64
         |--S. tinvenkiangG31
         |--S. trivialis Xia in Ding, Xia et al. 1984DX84
         |--S. tschernyschewiCB86
         |--S. ulanqabensis Xia in Ding, Xia et al. 1984DX84
         |--S. wongwenhaoG31
         `--S. zhesiensis Xia in Ding, Xia et al. 1984DX84

Sphaeroschwagerina sphaerica Scherbovich 1949 [incl. Schwagerina princeps Schellwien 1898 non Borelis princeps Ehrenberg 1842, *Sp. princeps]R-CB96

*Type species of generic name indicated


[CB86] Chen D.-Q. & Bao H. 1986. Lower Permian ostracodes from the Chihsia Formation of Jurong and Longtan, Jiangsu Province. Acta Micropalaeontologica Sinica 3 (2): 107–132.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3rd ed. Harvard University Press: Cambridge (Massachusetts).

[DX84] Ding Y., Xia G., Duan C., Li W., Liu X. & Liang Z. 1984. Study on the early Permian stratigraphy and fauna in Zhesi district, Nei Mongol Zizhiqu (Inner Mongolia). Bulletin Tianjin Institure Geol. Min. Res. 10.

[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[R-CB96] Rauzer-Chernousova, D. M., F. R. Bensh, M. V. Vdovenko, N. B. Gibshman, E. Y. Leven, O. A. Lipina, E. A. Reitlinger, M. N. Solovieva & I. O. Chedija. 1996. Spravočnik po Sistematike Foraminifer Paleozoâ (Èndotiroidy, Fuzulinoidy). Rossijskaâ Akademiâ Nauk, Geologičeskij Institut, Moskva “Nauka”.

[RR79] Ross, C. A., & J. R. P. Ross. 1979. Permian. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A291–A350. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

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