Scleria terrestris, copyright Mark Marathon.

Belongs within: Cyperaceae.

Scleria is a genus of sedges bearing unisexual flowers, with female flowers not surrounded by a perigynium. Members include S. pygmaea, a small annual (rarely more than five centimetres in height) found in northern Australia in which the two upper glumes of the female spikelet enclose the nut (Bentham 1878).

Characters (from Bentham 1878): Perennials or rarely annuals. Stems triquetrous, leafy, leaf-sheaths often but not always terminating in prominent ligula opposite lamina. Spikelets in clusters or small corymbose or oblong panicles, terminal and axillary, or in large compound terminal panicle. Flowers unisexual in unisexual or androgynous spikelets, female flowers always solitary in spikelet, males usually several, placed above female one in androgynous spikelets. Glumes imbricate all round, one to three outer ones empty. No hypogynous bristles or scales. Stamens 1 to 3. Style slender, deciduous, with three filiform stigmatic branches. Nut globular or ovoid, usually white, raised on a thickened entire or more or less three-lobed single or double disk.

<==Scleria [incl. S. ser. Subspicatae; Sclerieae]B78
|--S. browniiSB19 [incl. S. distansB78, S. pallidiflora Boeckel 1875B78]
`--+--+--S. lithospermaSB19 [incl. Hypoporum capillareB78, S. capillarisB78, H. sieberiB78]
| | |--S. l. var. lithospermaB78
| | `--S. l. var. linearisB78
| `--+--S. castaneaSB19
| `--+--S. pergracilisSB19
| `--+--S. bulbiferaSB19
| `--S. pantadeniaSB19
`--+--+--S. ciliataSB19
| |--S. oliganthaSB19
| `--S. triglomerataSB19
`--+--+--+--S. greigiifoliaSB19
| | `--S. ser. LaxaeB78
| | |--S. laxa [incl. S. novae-hollandiae Boeckel 1875]B78
| | `--S. rugosa Br. 1810SB19, G72 (see below for synonymy)
| `--+--S. lacustrisSB19
| `--S. poiformisSB19
`--+--+--S. melanomphalaSB19
| `--+--S. cyperinaSB19
| `--S. secansSB19
`--+--S. carphiformisSB19
`--+--+--S. bifloraSB19
| `--S. foliosaSB19
`--+--S. iostephanaSB19
`--+--+--S. ciliarisSB19
| `--+--S. levisSB19 [incl. S. zeylanicaG72]
| `--S. terrestrisSB19
`--+--+--S. flagellum-nigrorumSB19
| `--+--S. bracteataSB19
| `--S. latifoliaSB19
`--+--+--S. eggersianaSB19
| `--S. microcarpaSB19
`--+--S. mitisSB19
`--S. racemosaSB19

Scleria incertae sedis:
S. canescensSWK87
S. flagellumC55
S. lingulataLK14
S. mackaviensis Boeckel 1875B78
S. malaccensisB78
S. muehlenbergii Steudel 1841CRG03
S. paucifloraV72
S. polycarpaB00
S. psilorrhizaLK14
S. reflexaC55
S. reticularis Michaux 1803CRG03
|--S. r. var. reticularisCRG03
`--S. r. var. pubescensCRG03
S. scrobiculataB78
S. setoso-asperula Boeckel 1875B78
S. sumatrensisB00
S. thwaitesianaG72
S. verticillataV72
S. (ser. Axillares) pygmaea [=Diplacrum pygmaeum, Hypoporum pygmaeum, Sphaeropus pygmaeus]B78
S. (ser. Dioicae) sphacelataB78
S. (ser. Macrostachyae) oryzoidesB78
S. ser. PolystachyaeB78
|--S. chinensisB78
|--S. graeffeanaB78
|--S. hebecarpa [incl. S. dietrichiae Boeckel 1875]B78
|--S. margaritiferaB78
`--S. tesselata [incl. S. steudeliana, S. uliginosa]B78
|--S. t. var. tesselataB78
`--S. t. var. debilisB78

Scleria rugosa Br. 1810SB19, G72 [incl. S. flaccida Clarke 1894 non Steud. 1855G72, S. lateriflora Boeck. 1874G72]

*Type species of generic name indicated


[B78] Bentham, G. 1878. Flora Australiensis: A description of the plants of the Australian Territory vol. 7. Roxburghiaceae to Filices. L. Reeve & Co.: London.

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).

[CRG03] Camelbeke, K., A. A. Reznicek & P. Goetghebeur. 2003. (1850) Proposal to conserve the name Scleria reticularis with a conserved type (Cyperaceae). Taxon 52: 355–356.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[G72] Govindarajalu, E. 1972. Studies in Cyperaceae IV. Notes on Scleria rugosa R. Br. and its complex. Journal of the Bombay Natural History Society 69 (1): 246–249.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[SB19] Semmouri, I., K. Bauters, É. Léveillé-Bourret, J. R. Starr, P. Goetghebeur & I. Larridon. 2019. Phylogeny and systematics of Cyperaceae, the evolution and importance of embryo morphology. Botanical Review 85: 1–39.

[SWK87] Snyder, N. F. R., J. W. Wiley & C. B. Kepler. 1987. The Parrots of Luquillo: Natural history and conservation of the Puerto Rican parrot. Western Foundation of Vertebrate Zoology: Los Angeles.

[V72] Voss, E. G. 1972. Michigan Flora. Part I. Gymnosperms and Monocots. Cranbrook Institute of Science and University of Michigan Herbarium.

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