Sphex

 Great black wasp Sphex pensylvanicus, copyright Rodney Campbell.

Belongs within: Sphecidae.

Sphex is a cosmopolitan genus of medium-sized to very large, ground-nesting wasps that provision their nests with various Orthoptera. The majority of species are included in the subgenus Sphex, with the smaller subgenus Fernaldina differing in the absence of cerci and of a spiracular groove on the propodeum (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Male antenna usually with one or more non-spiculate, flat (rarely curved) placoids whilch are most common on flagellomeres IV–VI; free margin of female clypeus usually arcuate, sharp-edged, often with a small median notch, sometimes with a small median lobe which may be notched; clypeal margin usually broadly truncate in male, sometimes thickened, often broadly, shallowly emarginate; labrum in female usually with a median longitudinal carina which may project apically, surface plain in male; female mandible long, broadest at middle, usually strongly arcuate, the axes of the base and apex approximating a right angle, apex usually attaining or surpassing base of opposite mandible when folded, inner margin with single, large tooth near middle, the upper surface of which is usually ridged and sulcate, proximal side of inner tooth often subtended by a small tooth; male mandible shorter and more slender than female, inner tooth sometimes subapical; malar space absent; collar with some thickness and usually standing apart from scutum; propodeal enclosure present or absent; propodeal side usually with a complete spiracular groove; basal veinlet of submarginal cell II usually less than or equal to length of anterior veinlet, occasionally up to 1.2 times as long as anterior veinlet, anterior veinlct of submarginal cell III (part of radial sector) usually shorter than posterobasal veinlet (part of media); outer margin of female foreleg with well-developed rake comprised of long, often blade-like setae, tarsomere I usually asymmetrical and bearing rake setae on both outer and inner margins, II with two or more outer rake setae in addition to those at outer apex; apicoventral blade-like setae of last tarsomere of mid and hind legs narrow to broad, usually separated by about 1.5–2.0 times breadth of one seta, rarely separated by as much as 3.0–4.0 times a setal width; claws short to long, moderately to strongly arcuate in lateral profile, not prehensile, claw teeth perpendicularly or obliquely oriented to inner margin; petiole usually straight, short, length measured along dorsum rarely more than two-thirds combined length of hindtarsomeres II–IV; tergum VIII of male with cerci; sterna of male with or without long erect setae, transverse bands of fimbriae absent, terminal sterna often with tufts or patches of short, dense setae; sternum VIII usually triangular, or rounded and with a small apicomedian projection, but unusually modified in a few species; ventral margin of penis valve head often with teeth which usually extend in a row around outer surface near apex, penis valve head sometimes inflated and/or bearing apicodorsal or lateral “wings” or other processes; digitus of volsella simple except in some species with highly modified aedeagus.

<==Sphex Linnaeus 1758C85 [=Sphaex Scopoli 1772BM76]
    |--S. (Sphex)C85 [incl. Ammobia Billberg 1820BM76, Proterosphex Fernald 1905BM76]
    |    |--*S. (S.) flavipennis (Fabricius 1793)C85 (see below for synonymy)
    |    |--S. (S.) abyssinicus (Arnold 1928) [=Chlorion abyssinicum]BM76
    |    |--S. (S.) afer Lepeletier 1845BM76
    |    |    |--S. a. aferBM76
    |    |    `--S. a. sordidus Dahlbom 1845 (see below for synonymy)BM76
    |    |--S. (S.) ahasverus Kohl 1890C85
    |    |--S. (S.) alacer Kohl 1895BM76
    |    |--S. (S.) argentatus Fabricius 1787C85 (see below for synonymy)
    |    |    |--S. a. argentatusBM76
    |    |    `--S. a. fumosus Kohl 1890BM76
    |    |--S. (S.) argentinus Taschenberg 1869BM76
    |    |--S. (S.) ashmeadi (Fernald 1906) [=Chlorion ashmeadi]BM76
    |    |--S. (S.) atropilosus Kohl 1885 [incl. S. atrohirtus Kohl 1890]BM76
    |    |--S. (S.) basilicus (Turner 1915) [=Chlorion (Proterosphex) basilicus]C85
    |    |--S. (S.) bilobatus Kohl 1895 [incl. S. canescens Smith 1856 non Scopoli 1786]C85
    |    |--S. (S.) bohemanni Dahlbom 1845 (see below for synonymy)BM76
    |    |--S. (S.) brachystomus Kohl 1890BM76
    |    |--S. (S.) brasilianus Saussure 1867BM76
    |    |--S. (S.) caeruleanus Drury 1773 [incl. S. pulchripennis Mocsáry 1883]BM76
    |    |--S. (S.) caliginosus Erichson 1848 (see below for synonymy)BM76
    |    |--S. (S.) camposi Campos 1922BM76
    |    |--S. (S.) carbonicolor van der Vecht 1973 [=S. carbonaria Smith 1856 non Scopoli 1763]C85
    |    |--S. (S.) castaneipes Dahlbom 1843BM76
    |    |--S. (S.) cognatus Smith 1856 (see below for synonymy)C85
    |    |--S. (S.) confrater Kohl 1890 [incl. S. sieberti Strand 1910]BM76
    |    |--S. (S.) cubensis (Fernald 1906) [=Chlorion cubense; incl. S. clavipes Kohl 1890 non Linnaeus 1758]BM76
    |    |--S. (S.) darwiniensis Turner 1912C85
    |    |--S. (S.) decipiens Kohl 1895BM76
    |    |    |--S. d. decipiensBM76
    |    |    `--S. d. meridionalis (Arnold 1947) [=Chlorion meridionale]BM76
    |    |--S. (S.) decoratus Smith 1873 [=S. decorata]C85
    |    |--S. (S.) deplanatus Kohl 1895BM76
    |    |--S. (S.) diabolicus Smith 1858BM76
    |    |    |--S. d. diabolicusBM76
    |    |    `--S. d. flammitrichus Strand 1913 [incl. S. aureopilosus Berland 1928]BM76
    |    |--S. (S.) dorsalis Lepeletier 1845 (see below for synonymy)BM76
    |    |--S. (S.) dorycus Guérin-Méneville 1838 [incl. S. errabundus Kohl 1898]BM76
    |    |--S. (S.) ephippium Smith 1856C85
    |    |--S. (S.) ermineus Kohl 1890C85
    |    |--S. (S.) erythrinus Guiglia 1938 [=S. erytrhinus Magretti 1905 (n. n.)]BM76
    |    |--S. (S.) ferrugineipes Fox 1897BM76
    |    |--S. (S.) finschii Kohl 1890C85
    |    |--S. (S.) flavovestitus Smith 1856 (see below for synonymy)BM76
    |    |    |--S. f. flavovestitusBM76
    |    |    `--S. f. saussurei (Fernald 1906) [=Chlorion saussurei; incl. S. hirsutus Saussure 1867 non Scopoli 1763]BM76
    |    |--S. (S.) formosellus van der Vecht 1957C85
    |    |--S. (S.) fulvohirtus Bingham 1890BM76
    |    |--S. (S.) fumicatus Christ 1791 (see below for synonymy)BM76
    |    |    |--S. f. fumicatusBM76
    |    |    `--S. f. voeltzkowii Kohl 1909BM76
    |    |--S. (S.) fumipennis Smith 1856C85
    |    |    |--S. f. fumipennisBM76
    |    |    |--S. f. antennatus Smith 1856BM76
    |    |    `--S. f. rouxi Schulthess 1915 [incl. S. splendidus Berland 1928 nec Müller 1776 nec Reich 1795]BM76
    |    |--S. (S.) funerarius Gussakovskij 1934BM76
    |    |--S. (S.) gaullei Berland 1927BM76
    |    |--S. (S.) gilberti Turner 1908C85
    |    |--S. (S.) gisteli Strand 1917 (see below for synonymy)BM76
    |    |--S. (S.) gratiosus Smith 1856BM76
    |    |--S. (S.) guatemalensis Cameron 1888BM76
    |    |--S. (S.) habenus Say 1832 (see below for synonymy)BM76
    |    |--S. (S.) haemorrhoidalis Fabricius 1781 [incl. S. pachydermus Strand 1916, S. volubilis Kohl 1895]BM76
    |    |    |--S. h. haemorrhoidalisBM76
    |    |    |--S. h. basuto (Arnold 1947) [=Chlorion basuto]BM76
    |    |    |--S. h. kobrowi (Arnold 1928) [=Chlorion kobrowi]BM76
    |    |    |--S. h. mweruensis (Arnold 1947) [=Chlorion mweruense]BM76
    |    |    `--S. h. umtalicus Strand 1916BM76
    |    |--S. (S.) ichneumoneus (Linnaeus 1758) (see below for synonymy)BM76
    |    |--S. (S.) incomptus Gerstaecker 1871 [incl. Chlorion nyanzae Turner 1918]BM76
    |    |    |--S. i. incomptusBM76
    |    |    `--S. i. anonymus Leclercq 1955BM76
    |    |--S. (S.) ingens Smith 1856BM76
    |    |--S. (S.) inusitatus Yasumatsu 1935BM76
    |    |    |--S. i. inusitatusBM76
    |    |    `--S. i. fukuiensis Tsuneki 1957BM76
    |    |--S. (S.) jamaicensis (Drury 1773) (see below for synonymy)BM76
    |    |--S. (S.) jansei Cameron 1910BM76
    |    |--S. (S.) kolthoffi Gussakovskij 1938BM76
    |    |--S. (S.) lanatus Mocsáry 1883BM76
    |    |--S. (S.) latreillei Lepeletier 1831 [incl. S. chiliensis Lepeletier 1845, S. thunbergi Lepeletier 1831]BM76
    |    |--S. (S.) latro Erichson 1848 [incl. S. clypeatus Smith 1856, S. roratus Kohl 1890]BM76
    |    |--S. (S.) libycus Beaumont 1956BM76
    |    |--S. (S.) luctuosus Smith 1856 [=S. luctuosa]C85
    |    |--S. (S.) madasummae van der Vecht 1973 (see below for synonymy)BM76
    |    |--S. (S.) malagassus Saussure 1890BM76
    |    |--S. (S.) mandibularis Cresson 1868BM76
    |    |--S. (S.) maximiliani Kohl 1890BM76
    |    |--S. (S.) melanopus Dahlbom 1843 (see below for synonymy)BM76
    |    |--S. (S.) melas Gussakovskij 1930BM76
    |    |--S. (S.) mendozanus Brèthes 1909BM76
    |    |--S. (S.) mimulus Turner 1910C85
    |    |--S. (S.) mochii Giordani Soika 1942BM76
    |    |--S. (S.) modestus Smith 1856 [=S. modesta; incl. S. bannitus Kohl 1895, S. dolichocerus Kohl 1890]C85
    |    |--S. (S.) muticus Kohl 1885BM76
    |    |--S. (S.) neavei (Arnold 1928) [=Chlorion neavei]BM76
    |    |--S. (S.) nigrohirtus Kohl 1895 [incl. S. camerunicus Strand 1916, S. conradti Berland 1927]BM76
    |    |--S. (S.) nitidiventris Spinola 1851 [incl. S. beatus Cameron 1888, S. neotropicus Kohl 1890]BM76
    |    |--S. (S.) nudus Fernald 1903 [incl. S. bridwelli Fernald 1903]BM76
    |    |--S. (S.) obscurus (Fabricius 1804) (see below for synonymy)BM76
    |    |--S. (S.) observabilis (Turner 1918) [=Chlorion observabile]BM76
    |    |--S. (S.) opacus Dahlbom 1845 [incl. S. iheringii Kohl 1890]BM76
    |    |--S. (S.) optimus Smith 1856BM76
    |    |--S. (S.) oxianus Gussakovskij 1928BM76
    |    |    |--S. o. oxianusBM76
    |    |    `--S. o. nubilus Beaumont 1968BM76
    |    |--S. (S.) paulinierii Guérin-Méneville 1843 [incl. S. eximius Lepeletier 1845]BM76
    |    |--S. (S.) pensylvanicus Linnaeus 1763 [incl. S. robustisoma Strand 1916]BM76
    |    |--S. (S.) permagnus (Willink 1951) [=Chlorion permagnum]BM76
    |    |--S. (S.) peruanus Kohl 1890BM76
    |    |--S. (S.) praedator Smith 1859 (see below for synonymy)BM76
    |    |    |--S. p. praedatorBM76
    |    |    |--S. p. luteipennis Mocsáry 1883BM76
    |    |    |--S. p. melanopoda Strand 1915BM76
    |    |    `--S. p. tyrannicus Smith 1860BM76
    |    |--S. (S.) prosper Kohl 1890BM76
    |    |--S. (S.) pruinosus Germar 1817 (see below for synonymy)BM76
    |    |--S. (S.) resinipes (Fernald 1906) [=Chlorion resinipes; incl. S. rufipes Lepeletier 1845 non Linnaeus 1758]BM76
    |    |--S. (S.) resplendens Kohl 1885C85 (see below for synonymy)
    |    |--S. (S.) rhodosoma (Turner 1915) [=Chlorion (Proterosphex) rhodosoma]C85
    |    |--S. (S.) rufinervis Pérez 1895BM76
    |    |--S. (S.) rufiscutis (Turner 1918) [=Chlorion rufiscute]BM76
    |    |    |--S. r. rufiscutisBM76
    |    |    `--S. r. ‘laevigatus’ Arnold 1951 non Rossi 1794BM76
    |    |--S. (S.) rufocinctus Brullé 1833 (see below for synonymy)BM76
    |    |    |--S. r. rufocinctusBM76
    |    |    `--S. r. mavromoustakisi Beaumont 1947BM76
    |    |--S. (S.) rugifer Kohl 1890C85
    |    |--S. (S.) satanas Kohl 1898 [incl. S. gorgon Kohl 1913]BM76
    |    |--S. (S.) schoutedeni Kohl 1913BM76
    |    |--S. (S.) schrottkyi (Bertoni 1918) [=Proterosphex schrottkyi; incl. S. luciati Brèthes 1918]BM76
    |    |--S. (S.) semifossulatus van der Vecht 1973 [=S. argentifrons Smith 1868 non Lepeletier 1845]C85
    |    |--S. (S.) sericeus (Fabricius 1804)C85 [=Pepsis sericeaBM76]
    |    |    |--S. s. sericeusC85
    |    |    |--S. s. fabricii Dahlbom 1843 (see below for synonymy)BM76
    |    |    |--S. s. ferocior van der Vecht & Krombein 1955 [incl. S. ferox Smith 1862 non Chlorion ferox Westwood 1837]BM76
    |    |    |--S. s. godeffroyi Saussure 1869C85 [incl. S. aurifex Smith 1873C85, S. pallidehirtus Kohl 1890BM76]
    |    |    |--S. s. lineolus Lepeletier 1845 [incl. S. lepeletierii Saussure 1867, S. rugosus Matsumura 1912]BM76
    |    |    |--S. s. nigrescens van der Vecht & Krombein 1955BM76
    |    |    |--S. s. stueberi van der Vecht & Krombein 1955BM76
    |    |    `--S. s. wegneri van der Vecht & Krombein 1955BM76
    |    |--S. (S.) servillei Lepeletier 1845 (see below for synonymy)BM76
    |    |--S. (S.) stadelmanni Kohl 1895BM76
    |    |    |--S. s. stadelmanniBM76
    |    |    `--S. s. integrus (Arnold 1928) [=Chlorion integrum]BM76
    |    |--S. (S.) staudingeri Gribodo 1894C85
    |    |--S. (S.) subhyalinus Fox 1899BM76
    |    |--S. (S.) subtruncatus Dahlbom 1843 (see below for synonymy)BM76
    |    |    |--S. s. subtruncatusBM76
    |    |    |--S. s. coraxus van der Vecht 1957BM76
    |    |    |--S. s. orius van der Vecht 1957BM76
    |    |    |--S. s. siamensis Taschenberg 1869BM76
    |    |    |--S. s. sulciscuta Gribodo 1894BM76
    |    |    `--S. s. xuthus van der Vecht 1957BM76
    |    |--S. (S.) taschenbergi Magretti 1884BM76
    |    |--S. (S.) tepanecus Saussure 1867 [incl. S. mexicanus Taschenberg 1869 non Saussure 1867]BM76
    |    |--S. (S.) texanus Cresson 1872BM76
    |    |--S. (S.) tinctipennis Cameron 1888BM76
    |    |--S. (S.) tomentosus Fabricius 1787 [incl. S. tuberculatus Smith 1873 non Villers 1789]BM76
    |    |--S. (S.) torridus Smith 1873BM76
    |    `--S. (S.) vestitus Smith 1856 [=S. vestita; incl. S. imperialis Kohl 1890, S. praetexta Smith 1873]C85
    `--S. (Fernaldina Bohart & Menke 1963)BM76
         |--S. (*F.) lucae Saussure 1867 [incl. S. belfragei Cresson 1872]BM76
         `--S. (F.) melanocnemis Kohl 1885BM76
Sphex incertae sedis:
  S. albifrons Villers 1789BM76
  S. annularis Poda 1761BM76
  S. antarcticus Linnaeus 1767BM76
  S. argillacea Linnaeus 1758L58
  S. aurata Linnaeus 1758L58
  S. austriacus Schrank 1781BM76
  S. bifasciatus Müller 1776BM76
  S. bischoffi Zeuner 1931BM76
  S. caerulea Linnaeus 1758L58
  S. cinctus’ Fabricius 1793 non S. cincta Scopoli 1763BM76
  S. coccineus Gmelin 1790BM76
  S. collaris Linnaeus 1767BM76
  S. colon Linnaeus 1758L58
  S. conicus Villers 1789BM76
  S. ‘conicus’ Radoszkowski 1877 non Villers 1789BM76
  S. cyanea Linnaeus 1758L58
  S. elongatus Villers 1789BM76
  S. fabulosa Linnaeus 1758L58
  S. ‘flavicornis’ Villers 1789 non Fabricius 1781BM76
  S. gibbosus Rossi 1790BM76
  S. giganteus Heer 1867BM76
  S. gregarius Scopoli 1763BM76
  S. guttatus Gmelin 1790BM76
  S. ignita Linnaeus 1758L58
  S. indica Linnaeus 1758L58
  S. leucopterus Pallas 1771BM76
  S. lineatus Villers 1789BM76
  S. lucidus Villers 1789BM76
  S. lugubris Villers 1789BM76
  S. mauritanicus Linnaeus 1767 [incl. S. mauritianus Christ 1791]BM76
  S. melanochlorus Gmelin 1790BM76
  S. militaris Müller 1776BM76
  S. mixtus Fabricius 1794BM76
  S. naviculatus Villers 1789BM76
  S. ‘niger’ Müller 1775 non Fabricius 1775BM76
  S. ‘nigerrimus’ Schrank 1802 non Scopoli 1763BM76
  S. nigrita Lucas 1849E12
  S. ‘obscurus’ Statz 1936 nec Pepsis obscura Fabricius 1804 nec S. obscurus Fischer-Waldheim 1843BM76
  S. pectinipes Linnaeus 1758L58
  S. pygmaeus Schrank 1785BM76
  S. quadrifasciatus Müller 1776BM76
  S. ‘quadrifasciatus’ Villers 1789 non Müller 1776BM76
  S. semicinctus Villers 1789BM76
  S. serotinus Müller 1776BM76
  S. spinipes Gmelin 1790BM76
  S. splendidus Müller 1776BM76
  S. ‘splendidus’ Reich 1795 non Müller 1776BM76
  S. stigma Linnaeus 1767BM76
  S. tanoi Tsuneki 1974BM76
  S. tenthredinoides Scopoli 1763BM76
  S. testaceus Gmelin 1790BM76
  S. ‘tomentosus’ Gmelin 1790 non Fabricius 1787BM76
  S. triangulum Villers 1789 [=S. triangulus (l. c.)]BM76
  S. tricolor’ Reich 1795 nec Schrank 1781 nec Fabricius 1793BM76
  S. trifasciatus Müller 1776BM76
  S. trimarginatus Müller 1776BM76
  S. tropica Linnaeus 1758L58
  S. vaga Linnaeus 1758L58
  S. variegata Linnaeus 1758L58
  S. ‘vespiformis’ Schrank 1781 non Fabricius 1775BM76
Nomina nuda: S. annulatus Lichtenstein 1796 non S. annulata Fabricius 1798BM76
             S. antaeus Lichtenstein 1796BM76
             S. argyrosticta Lichtenstein 1796BM76
             S. argyrostoma Lichtenstein 1796BM76
             S. aselenos Lichtenstein 1796BM76
             S. aureonitens Lichtenstein 1796BM76
             S. brachyptera Lichtenstein 1796BM76
             S. chilon Lichtenstein 1796BM76
             S. chrysosticta Lichtenstein 1796BM76
             S. chrysostoma Lichtenstein 1796BM76
             S. corynetes Lichtenstein 1796BM76
             S. formicaroides Gistel 1837BM76
             S. gigas Lichtenstein 1796BM76
             S. gnatho Lichtenstein 1796BM76
             S. lineatus Lichtenstein 1796 non Villers 1789BM76
             S. manduco Lichtenstein 1796BM76
             S. manicatus Lichtenstein 1796BM76
             S. mesomelaena Lichtenstein 1796BM76
             S. nigellus Lichtenstein 1796 non Smith 1856BM76
             S. nigripes Lichtenstein 1796 nec Fabricius 1793 nec Smith 1856BM76
             S. pumila Lichtenstein 1796BM76
             S. quinqueguttatus Lichtenstein 1796BM76
             S. ruficollis Lichtenstein 1796BM76
             S. rufilumbis Lichtenstein 1796BM76
             S. serpentinus Lichtenstein 1796BM76
             S. sinensis Lichtenstein 1796BM76
             S. synoecodes Perez in Campos 1922BM76
             S. tiphia Gistel 1837BM76

Sphex (Sphex) afer sordidus Dahlbom 1845 [incl. S. pachysoma Kohl 1890, S. plumipes Radoszkowski 1886 non Drury 1773, S. tristis Kohl 1885]BM76

Sphex (Sphex) argentatus Fabricius 1787C85 [=Pepsis argentataBM76, *Ammobia argentataBM76; incl. S. argenteus Turton 1802BM76, S. argentifrons Lepeletier 1845BM76, S. nanulus Strand 1913BM76, S. plumiferus Costa 1864BM76, S. umbrosus Christ 1791C85, S. unicolor Fabricius 1787BM76]

Sphex (Sphex) bohemanni Dahlbom 1845 [incl. S. abbottii Fox 1891, S. kilimandjaroensis Cameron 1908, S. transvaalensis Cameron 1910]BM76

Sphex (Sphex) caliginosus Erichson 1848 [incl. S. erythropterus Cameron 1888, S. fuscus Lepeletier 1845 non Linnaeus 1761]BM76

Sphex (Sphex) cognatus Smith 1856 [=S. cognata; incl. S. amator Smith 1856, S. formosa Smith 1856, S. opulenta Smith 1856]C85

Sphex (Sphex) dorsalis Lepeletier 1845 [incl. S. chlorargyricus Costa 1862, S. dubitatus Cresson 1872, S. micans Taschenberg 1869 non Eversmann 1849, S. singularis Smith 1856, S. spiniger Kohl 1890]BM76

*Sphex (Sphex) flavipennis (Fabricius 1793)C85 [=Pepsis flavipennisC85; incl. S. bicolor Dahlbom 1845 non Fabricius 1775BM76, S. cinereorufocinctus Dahlbom 1845BM76, S. rufodorsatus De Stefani 1887BM76, S. sellae Gribodo 1873BM76]

Sphex (Sphex) flavovestitus Smith 1856 [incl. S. flavipes Smith 1856 nec Fabricius 1781 nec Christ 1791, Chlorion flavitarse Fernald 1906]BM76

Sphex (Sphex) fumicatus Christ 1791 [incl. S. albifrons Fabricius 1793 non Villers 1789, S. argentiferus Walker 1871, Chlorion davisi Fernald 1907, S. erebus Kirby 1900, S. magrettii Gribodo 1894, S. metallicus Taschenberg 1869]BM76

Sphex (Sphex) gisteli Strand 1917 [incl. S. aurulentus Gistel 1857 nec Fabricius 1787 nec Fabricius 1793 nec Guérin-Méneville 1835]BM76

Sphex (Sphex) habenus Say 1832 [incl. S. chrysophorus Kohl 1890, S. illustris Cresson 1872, S. lanciger Kohl 1895, S. lautus Cresson 1872, S. princeps Kohl 1890]BM76

Sphex (Sphex) ichneumoneus (Linnaeus 1758) [=Apis ichneumonea; incl. S. aurifluus Perty 1833, S. aurocapillus Templeton 1841, S. croesus Lepeletier 1845, S. dimidiatus Lepeletier 1845 nec De Geer 1773 nec Christ 1791 nec S. dimidiata Fabricius 1793, S. ignotus Strand 1916, S. sumptuosus Costa 1862, Nomada surinamensis Retzius 1783]BM76

Sphex (Sphex) jamaicensis (Drury 1773) [=Vespa jamaicensis; incl. S. aurulentus Guérin-Méneville 1835 nec Fabricius 1787 nec Fabricius 1793, S. fulviventris Kohl 1890, S. jamaica Christ 1791, S. lanierii Guérin-Méneville 1844, S. ornatus Lepeletier 1845]BM76

Sphex (Sphex) madasummae van der Vecht 1973 [incl. S. caerulescens Le Guillou 1841 non Reich 1795, S. maurus Smith 1856 non Fabricius 1787, S. nigerrimus Costa 1864 nec Scopoli 1763 nec Schrank 1802]BM76

Sphex (Sphex) melanopus Dahlbom 1843 [incl. S. difficilis Spinola 1851, S. funestus Kohl 1890, S. proximus Smith 1856, S. ruficaudus Taschenberg 1869]BM76

Sphex (Sphex) obscurus (Fabricius 1804) [=Pepsis obscura; incl. S. cinerascens Dahlbom 1843, S. hirtipes Fabricius 1793, S. xanthopterus Cameron 1889]BM76

Sphex (Sphex) praedator Smith 1859 [incl. S. calopterus Kohl 1890, S. celebesianus Strand 1913, S. kohlianus Strand 1913]BM76

Sphex (Sphex) pruinosus Germar 1817 [incl. S. retractus Nurse 1903, S. rothneyi Cameron 1889, S. scioensis Gribodo 1879, S. vicinus Lepeletier 1845]BM76

Sphex (Sphex) resplendens Kohl 1885C85 [=S. nitidiventris Smith 1859 non Spinola 1851C85; incl. S. gratiosissimus Dalla Torre 1897C85, S. gratiosa Smith 1859 non S. gratiosus Smith 1856C85, S. lanceiventris Vachal 1908BM76, S. mertoni Strand 1911BM76, S. wallacei Turner 1908C85]

Sphex (Sphex) rufocinctus Brullé 1833 [incl. S. leuconotus Brullé 1833, S. maxillosus Fabricius 1793 non Poiret 1787, *Proterosphex maxillosus, S. obscurus Fischer-Waldheim 1843 non Pepsis obscura Fabricius 1804, S. pedibusnigris Zanon 1925, S. triangulum Brullé 1833 non Villers 1789]BM76

Sphex (Sphex) sericeus fabricii Dahlbom 1843 [incl. S. aurulentus Fabricius 1793 non Fabricius 1787, S. ferrugineus Lepeletier 1845]BM76

Sphex (Sphex) servillei Lepeletier 1845 [incl. S. chichimecus Saussure 1867, S. congener Kohl 1890, S. fuliginosus Dahlbom 1843 non Scopoli 1763, S. joergenseni Brèthes 1913]BM76

Sphex (Sphex) subtruncatus Dahlbom 1843 [incl. S. erythropoda Cameron 1889, S. nigripes Smith 1856 non Fabricius 1793, S. tsingtauensis Strand 1916]BM76

*Type species of generic name indicated

References

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

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