Laemostenus complanatus, copyright Udo Schmidt.

Belongs within: Platynini.

The Sphodrina are a primarily Palaearctic group of ground beetles characterised by features of the female genitalia: the gonostyli lack a fringe of setae on the penultimate segment, have the dorsal seta on the apical segment moved to the inner lateral margin, and possess preapical sensory setae on the apical segment.

Sphodrina [Sphodrini]
|--Sphodrus planus [=Harpalus (Sphodrus) planus]G20
|--Pristonychus Dejean 1828R-JC01, LM87 [=Prystonychus (l. c.)LM87]
| |--*P. terricola (Herbst 1784)LM87, BB01 [=Carabus tericolaLM87, Laemostenus (Pristonychus) terricolaBB01]
| |--P. alticola Fairmaire 1889F89b
| |--P. angustatusF89b
| |--P. barbarus Lucas 1846E12
| |--P. complanatusR-JC01
| |--P. davidis Fairmaire 1889F89a
| |--P. ghilianiiF89a
| |--P. oblongusF89b
| |--P. sardous Lucas 1846E12
| `--P. sulcipennis Fairmaire 1889F89a
`--Laemostenus Bonelli 1810E04, LM87 [=Laemosthenus (l. c.)LM87]
|--*L. venustus (Dejean 1828) [=Carabus venustus]LM87
|--L. cavicola [=Antisphodrus cavicola]PN02
| |--L. c. cavicolaPN02
| `--L. c. aeacus (Miller 1861)PN02
`--L. complanatus (Dejean 1828) [=Pristonychus complanatus; incl. P. australis Blackburn 1888]LM87

*Type species of generic name indicated


[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[E04] Emberson, R. M. 2004. A European ground beetle, Harpalus tardus (Panzer) (Coleoptera: Carabidae), in Canterbury, and the predisposition of European carabids to establish in exotic temperate locations. New Zealand Entomologist 27: 39–42.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F89a] Fairmaire, L. 1889a. Coléoptères de l’intérieur de la Chine. 5e partie. Annales de la Société Entomologique de France, 6e série 9: 5–84.

[F89b] Fairmaire, L. 1889b. Descriptions de deux carabiques du nord de l’Inde appartenant au Musée de Calcutta. Annales de la Société Entomologique de France, 6e série 9: xv–xvii.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[PN02] Poggi, R., G. Nonveiller, A. Colla, D. Pavićević & T. Rađa. 2001–2002. Thaumastocephalini, a new tribe of Pselaphinae for Thaumastocephalus folliculipalpus n. gen., n. sp., an interesting new troglobious species from central Dalmatia (Croatia) (Coleoptera, Staphylinidae, Pselaphinae). Annali del Museo Civico di Storia Naturale “Giacomo Doria” 94: 1–20.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

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