Diagram of axial section of Nankinella, from Loeblich & Tappan (1964).

Belongs within: Fusulinoida.

The Staffellidae are a group of smaller fusulinids known from the Upper Carboniferous and Permian (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Shell small, spherical to discoidal, with umbilicate or flush axial ends; septa strongly arched forward, closely spaced, and totally unfluted from end to end; tunnel singular, bordered by distinctly asymmetrical chomata except in terminal part of last volution; walls composed of upper and lower layers that are in turn underlain and overlain by secondary deposits in most forms, except for outer part of last volution.

<==Staffellidae [Staffellacea, Staffellinae, Staffellinidae, Staffellininae]LT64
    |--Nummulostegina Schubert 1907LT64
    |    `--*N. velebitana Schubert 1908LT64
    |--Eoverbeekina Lee 1933R-CB96
    |    `--*E. intermedia Lee 1933LT64
    |--Parastaffelloides Reitlinger 1963R-CB96
    |    `--*P. pseudosphaeroidea (Dutkevich 1934) [=Staffella pseudosphaeroidea]R-CB96
    |--Staffelloides Liem 1976R-CB96
    |    `--*S. polyaxica Liem 1976R-CB96
    |--Sphaerulina Lee 1933 non Sacc. 1878 (ICBN)LT64
    |    |--*S. crassispira Lee 1933LT64
    |    `--S. ogbinensisK04
    |--Praemisellina Kalmykova 1972R-CB96
    |    |--*P. georgii Kalmykova 1972R-CB96
    |    `--‘Staffella’ dagmarae Dutkevich 1934R-CB96
    `--Staffella Ozawa 1925LT64
         |--*S. sphaerica (Abich 1859) [=Fusulina sphaerica, Fusulinella sphaerica]LT64
         |--S. expansaLT64
         `--S. moellerana Thompson 1935LT64

*Type species of generic name indicated


[K04] Kobayashi, F. 2004. Late Permian foraminifers from the Limestone Block in the Southern Chichibu Terrane of west Shikoku, SW Japan. Journal of Paleontology 78 (1): 62–70.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[R-CB96] Rauzer-Chernousova, D. M., F. R. Bensh, M. V. Vdovenko, N. B. Gibshman, E. Y. Leven, O. A. Lipina, E. A. Reitlinger, M. N. Solovieva & I. O. Chedija. 1996. Spravočnik po Sistematike Foraminifer Paleozoâ (Èndotiroidy, Fuzulinoidy). Rossijskaâ Akademiâ Nauk, Geologičeskij Institut, Moskva “Nauka”.

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