Steganacarus

Steganacarus magnus, from Micropics.

Belongs within: Phthiracaridae.

Steganacarus is a genus of phthiracarid mites found in the Holarctic, Afrotropical and Neotropical regions in which seta d on tibia IV is long and removed from the solenidion.

Rhacaplacarus is a cosmopolitan genus of phthiracarid mites. The type species R. amoenus was described from Uganda and probably the greater number of known species are from Africa (Subías 2004).

<==Steganacarus Ewing 1917S04
    |--S. (Steganacarus) [incl. Trachyhoplophora Berlese 1923]S04
    |    |--S. (S.) magnus (Nicolet 1855)S04 (see below for synonymy)
    |    |--S. (S.) applicatus (Sellnick 1920)S04 (see below for synonymy)
    |    |--S. (S.) balearicus Pérez-Íñigo 1969S04
    |    |--S. (S.) bicarinatus Jeleva 1970S04
    |    |--S. (S.) caelestis Niedbała 1984S04
    |    |--S. (S.) carlosi Niedbała 1984S04
    |    |--S. (S.) carusoi Bernini & Avanzati 1989 [incl. S. applicatus crassisetosus Willmann 1939 (n. d.)]S04
    |    |--S. (S.) coniunctus Niedbała 1983 [incl. S. danae Mahunka & Miko 1989]S04
    |    |--S. (S.) donatoi Avanzati, Baratti & Bernini 1994S04
    |    |--S. (S.) doumandji Niedbała 1986S04
    |    |--S. (S.) flagellatissimus Mahunka 1979S04
    |    |--S. (S.) grandjeani Mahunka 1977 [incl. S. sacculiferus Parry 1978]S04
    |    |--S. (S.) guanarteme Pérez-Íñigo & Peña 1996S04
    |    |--S. (S.) herculeanus Willmann 1953 [incl. S. longisetosus Moraza 1985]S04
    |    |--S. (S.) hirsutus Pérez-Íñigo 1974S04
    |    |    |--S. h. hirsutus [incl. S. perezinigoi Balogh & Mahunka 1979]S04
    |    |    `--S. h. azorensis Pérez-Íñigo 1992S04
    |    |--S. (S.) inaestimabilis Niedbała 1984S04
    |    |--S. (S.) incognitus Niedbała 1984S04
    |    |--S. (S.) incomptus Niedbała 1983S04
    |    |--S. (S.) insulanus Pérez-Íñigo & Pérez-Íñigo 1996S04
    |    |--S. (S.) inurbanus Niedbała 1983S04
    |    |--S. (S.) manganellii Avanzati, Baratti & Bernini 1994S04
    |    |--S. (S.) maximus (Haupt 1882) (n. d.) [=Hoplophora maxima]S04
    |    |--S. (S.) michaeli Bernini & Avanzati 1987S04
    |    |--S. (S.) nemorosus Gordeeva 1991S04
    |    |--S. (S.) patruelis Niedbała 1983 [incl. S. lazitanicus Mahunka & Miko 1989]S04
    |    |--S. (S.) personatus Niedbała 1983S04
    |    |--S. (S.) similis Willmann 1939S04
    |    |--S. (S.) simonettae Bernini & Avanzati 1987S04
    |    |--S. (S.) sol Balogh 1958S04 [incl. Hoplophthiracarus peracutus Mahunka 1983S04, S. peracutusN86]
    |    |--S. (S.) spinosus (Sellnick 1920) (see below for synonymy)S04
    |    |--S. (S.) tenerifensis Pérez-Íñigo 1972S04
    |    |--S. (S.) travei Lions 1968S04
    |    |--S. (S.) vernaculus Niedbała 1982S04
    |    |--S. (S.) vestitus Niedbała 1983 [incl. S. complicatus Mahunka 1993]S04
    |    |--S. (S.) wallworki Mahunka 1984S04
    |    `--S. (S.) werneri Mahunka 1993S04
    |--‘Rhacaplacarus’ (Mantigueracarus Balogh & Mahunka 1992)S04
    |    |--R. (*M.) baggioi (Balogh & Mahunka 1992) [=*Mantigueracarus baggioi]S04
    |    |--R. (M.) aculeatus (Mahunka 1995) [=Hoplophthiracarus (Plonaphacarus) aculeatus]S04
    |    |--R. (M.) grandjeani (Niedbała 1987) [=Plonaphacarus grandjeani]S04
    |    |--R. (M.) incredibilis (Niedbała 1982)S04 [=Neophthiracarus incredibilisS04, Hoplophthiracarus incredibilisN86]
    |    `--R. (M.) perezinigoi (Balogh & Mahunka 1992) [=Mantigueracarus perezinigoi]S04
    |--S. (Rafacarus Niedbała 1982) [incl. Neosteganacarus Balogh & Mahunka 1992, Nortonacarus Balogh & Mahunka 1992]S04
    |    |--S. (*R.) rafalskii (Niedbała 1982)S04 (see below for synonymy)
    |    |--S. (R.) angulatus (Balogh & Mahunka 1992) [=Neosteganacarus angulatus]S04
    |    |--S. (R.) niedbalai (Pérez-Íñigo & Baggio 1996) [=Protophthiracarus niedbalai]S04
    |    |--S. (R.) relictus (Balogh & Mahunka 1992) [=Neosteganacarus relictus]S04
    |    `--S. (R.) valeriae (Balogh & Mahunka 1992)S04 [=*Nortonacarus valeriaeBM92, S04]
    |--S. (Rhacaplacarus Niedbała 1986)N06, S04
    |    |--S. (*R.) amoenus (Niedbała 1983)N86, S04 [=Hoplophorella amoenaS04]
    |    |--‘Rhacaplacarus’ (R.) ambiguus (Niedbała 1982) [=Hoplophthiracarus ambiguus]S04
    |    |--‘Rhacaplacarus’ (R.) brevisetus (Mahunka 1984) [=Hoplophthiracarus brevisetus]S04
    |    |--‘Rhacaplacarus’ (R.) contrarius (Niedbała 1984) [=Hoplophthiracarus contrarius]S04
    |    |--S. (R.) diaphoros Niedbała 2000S04
    |    |--‘Rhacaplacarus’ (R.) dikros (Niedbała 2000) [=Plonaphacarus dikros]S04
    |    |--‘Rhacaplacarus’ (R.) discrepus (Niedbała 1982) [=Hoplophthiracarus discrepus]S04
    |    |--‘Rhacaplacarus’ (R.) dispar (Niedbała 2000) [=Plonaphacarus dispar]S04
    |    |--‘Rhacaplacarus’ (R.) ecphylus (Niedbała 2001) [=Plonaphacarus ecphylus]S04
    |    |--‘Rhacaplacarus’ (R.) forsslundi (Niedbała 1987) [=Plonaphacarus forsslundi]S04
    |    |--‘Rhacaplacarus’ (R.) granulatus (Banks 1902) [=Hoploderma granulatum]S04
    |    |--S. (R.) inflatus (Niedbała 1984)N86, S04 [=Hoplophthiracarus inflatusS04]
    |    |--‘Rhacaplacarus’ (R.) insignitus (Niedbała 1989) [=Plonaphacarus insignitus]S04
    |    |--‘Rhacaplacarus’ (R.) insolens (Niedbała 2000) [=Plonaphacarus insolens]S04
    |    |--S. (R.) jacoti Niedbała 1987N06
    |    |--‘Rhacaplacarus’ (R.) kaszabi (Balogh 1988) [=Hoplophthiracarus kaszabi]S04
    |    |--S. (R.) laetabilis (Niedbała 1988)N86, S04 [=Hoplophorella laetabilisS04]
    |    |--‘Rhacaplacarus’ (R.) laterospinosus Mahunka 1993S04
    |    |--‘Rhacaplacarus’ (R.) lividus (Niedbała 2000) [=Hoplophthiracarus lividus]S04
    |    |--‘Rhacaplacarus’ (R.) loebli (Mahunka 1985) [=Hoplophthiracarus loebli]S04
    |    |--S. (R.) machadoi Balogh 1958S04 [=Plonaphacarus machadoiN86]
    |    |--‘Rhacaplacarus’ (R.) ngongi (Niedbała 2001) [=Plonaphacarus ngongi]S04
    |    |--‘Rhacaplacarus’ (R.) optivus (Niedbała 1994) [=Plonaphacarus optivus]S04
    |    |--S. (R.) ortizi Pérez-Íñigo 1970S04
    |    |--‘Rhacaplacarus’ (R.) persimilis (Niedbała 1994) [=Plonaphacarus persimilis]S04
    |    |--S. (R.) pervigens (Niedbała 1988)N86, S04 [=Hoplophorella pervigensS04]
    |    |--‘Rhacaplacarus’ (R.) rafalskii (Niedbała 1997) [=Plonaphacarus rafalskii]S04
    |    |--‘Rhacaplacarus’ (R.) sarawaki (Niedbała 2000) [=Austrophthiracarus sarawaki]S04
    |    |--‘Rhacaplacarus’ (R.) scrupeus (Niedbała 1989) [=Plonaphacarus scrupeus]S04
    |    |--S. (R.) spinatus Niedbała 2001S04
    |    |--‘Rhacaplacarus’ (R.) spiniferus Mahunka 1993S04
    |    |--S. (R.) spiniger (Aoki 1980)N86, S04 [=Hoplophorella spinigerS04]
    |    |--S. (R.) succinctus Niedbała 2001S04
    |    |--‘Rhacaplacarus’ (R.) tanzicus (Mahunka 1993) [=Hoplophthiracarus (Plonaphacarus) tanzicus]S04
    |    |--S. (R.) thoreaui Jacot 1930S04
    |    `--S. (R.) zicsii Mahunka 1988S04
    `--S. (Tropacarus Ewing 1917) [incl. Calhoplophora Berlese 1923]S04
         |--S. (*T.) carinatus (Koch 1841)S04 (see below for synonymy)
         |--S. (T.) balcanicus Berninni & Avanzati 1989S04
         |--S. (T.) boulfekhari Niedbała 1986S04
         |--S. (T.) brevipilus (Berlese 1923)S04 (see below for synonymy)
         |--S. (T.) callainii Bernini, Bernini & Avanzati 1989S04
         |--S. (T.) desmeti Niedbała 1988 [incl. S. (T.) altitudinis Bernini, Bernini & Avanzati 1989]S04
         |--S. (T.) excavatus (Niedbała 1981) [=Tropacarus excavatus]S04
         |--S. (T.) lasithiensis Mahunka 1979S04
         |--S. (T.) maghrebinus Bernini, Bernini & Avanzati 1989S04
         |--S. (T.) malatestai Avanzati, Baratti & Bernini 1994S04
         `--S. (T.) pseudocarinatus Bernini & Avanzati 1989S04

Steganacarus (*Rafacarus) rafalskii (Niedbała 1982)S04 [=*Rafacarus rafalskiiS04; incl. *Neosteganacarus cataracta Balogh & Mahunka 1992BM92, S04]

Steganacarus (Steganacarus) applicatus (Sellnick 1920)S04 [=Phthiracarus applicatusS04; incl. P. berlesei Oudemans 1915 non Michael 1898S04, Hoplophora quadriseriata Haupt 1882S04, Hoploderma quadriseriatumM98]

Steganacarus (Steganacarus) magnus (Nicolet 1855)S04 [=Hoplophorella magnaS04, Hoploderma magnumS22, Hoplophora magnaBA87; incl. Hoplophora anomala Berlese 1883S04, Hoploderma anomalaCH98, Phthiracarus anomalusBA88, *Steganacarus anomalusS04, P. magnus anomaloides Oudemans 1915S04, S. magnus f. corcyraea Bernini & Avanzati 1988BA88, S. karamani Tarman 1962S04, P. magna f. oblonga Oudemans 1915N86, P. magnus rotundus Oudemans 1915 non Ewing 1908S04, P. magnus transitans Oudemans 1915S04]

Steganacarus (Steganacarus) spinosus (Sellnick 1920) [=Phthiracarus spinosus; incl. S. barborae Mahunka & Miko 1989, S. punctulatus Sergienko 1985]S04

Steganacarus (Tropacarus) brevipilus (Berlese 1923)S04 [=Phthiracrus (Trachyhoplophora) brevipilusS04, Tropacarus curtipilus (l. c.)N86; incl. Hoploderma brevipilus perfecta Sellnick 1931S04]

Steganacarus (*Tropacarus) carinatus (Koch 1841)S04 [=Hoplophora carinataS04, Calhoplophora carinataN86, Hoploderma carinatumBA89, Hoplophora carinatumBA89; incl. Hoplophora carinata var. pulcherrima Berlese 1887N86, S04, Hoploderma pulcherrimumM98, Steganacarus (Tropacarus) pulcherrimusBA91]

*Type species of generic name indicated

References

[BM92] Balogh, J., & S. Mahunka. 1992. New phthiracarid taxa from Brazilian soils (Acari, Oribatida). Acta Zoologica Hungarica 38 (3–4): 159–174.

[BA87] Bernini, F., & A. M. Avanzati. 1987. Notulae oribatologicae XLI. Some new Steganacarus species belonging to the magnus group (Acarida, Oribatida). International Journal of Acarology 13 (3): 165–176.

[BA88] Bernini, F., & A. M. Avanzati. 1988. Taxonomic revision of Steganacarus (Steganacarus) magnus (Nicolet, 1855) (Acarida, Oribatida). Journal of Natural History 22: 435–464.

[BA89] Bernini, F., & A. M. Avanzati. 1989. Notulae oribatologicae XLVI. Steganacarus (Tropacarus) carinatus (C. L. Koch, 1841) and two new Mediterranean Tropacarus species (Acarida, Oribatida). Acarologia 30 (2): 143–161.

[BA91] Bernini, F., & A. M. Avanzati. 1991. A new approach to the systematics of the genus Steganacarus (Oribatida). In: Schuster, R., & P. W. Murphy (eds) The Acari: Reproduction, development and life-history strategies pp. 335–342. Chapman & Hall: London.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[M98] Michael, A. D. 1898. Oribatidae. In: H. Lohmann (ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen vol. 3. Acarina pp. 1–93. R. Friedländer und Sohn: Berlin.

[N86] Niedbała, W. 1986. Catalogue des Phthiracaroidea (Acari), clef pour la détermination des espèces et descriptions d’espèces nouvelles. Annales Zoologici 40 (4): 309–370.

[N06] Niedbała, W. 2006. Supplement to the knowledge of ptyctimous mites (Acari: Oribatida) from Australian region. Annales Zoologici 56 (Suppl. 1): 99–156.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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