Mounted skeleton of Simosthenurus occidentalis in the Victoria Fossil Cave, photographed by ccdoh1.

Belongs within: Macropodidae.

The Sthenurinae include the fossil short-faced kangaroos, some of which are among the largest known kangaroos. The late-surviving Procoptodon goliah reached approximately two metres in height and may have weighed over 200 kg. Sthenurines probably would have been browsers on leaves rather than grazers like modern kangaroos.

Characters (from Long et al. 2002): Skulls relatively short and deep, with wide rostra and thick ectotympanic bones. I1-3 tending to form narrow V-shaped arcade; I3 with elongate crown relative to I1 and I2. Lower jaws often fused at symphysis. Premolars usually wide, high-crowned, complex; molars with crenulated enamel, postlink crest on rear of upper molars; lower molars often with well-developed premetacristid blade extending forward to join paracristid. Digits I and IV on fore foot often greatly reduced, digits II and III elongate; hind foot usually with all digits except IV reduced or lost. Astragalus with high, rounded crests on tibial surface.

    |  i. s.: Rhizosthenurus Kear 2002BA12
    |           `--*R. flanneryi Kear 2002BA12
    |         ArchaeosimosBA12
    |           |--A. cegsai (Pledge 1992)BA12
    |           `--A. correlli Prideaux 2004BA12
    |--Wanburoo Cooke 1999PW10, LA02
    |    `--*W. hilarus Cooke 1999LA02
    `--+--Hadronomas Woodburne 1967LA02
       |    `--*H. puckridgi Woodburne 1967LA02
       `--+--Sthenurus Owen 1873LA02
          |    |  i. s.: S. notabilis Bartholomai 1963LA02
          |    |--S. andersoni Marcus 1962FS15, M62
          |    `--+--*S. atlas (Owen 1838)LA02, FS15, LA02 [=Macropus atlasF71]
          |       `--+--S. stirlingi Wells & Tedford 1995FS15, LA02
          |          `--S. tindalei Tedford 1966FS15, LA02
          `--+--Metasthenurus newtonae (Prideaux 2004)FS15, BA12
             `--+--Procoptodon Owen 1874LA02
                |    |  i. s.: P. oreasD07
                |    |         P. pusio Owen 1874LA02
                |    |         P. rapha Owen 1874LA02
                |    |         P. texasensis Archer 1978LA02
                |    |--P. browneorumFS15
                |    `--+--*P. goliah (Owen 1845)LA02, FS15, LA02 [=Macropus goliahF71]
                |       `--P. gilliFS15
                `--Simosthenurus Tedford 1966LA02
                     |--*S. occidentalis (Glauert 1910) [=Sthenurus (*Simosthenurus) occidentalis]LA02
                     |--S. antiquus (Bartholomai 1963)LA02 [=Sthenurus (Simosthenurus) antiquusF71]
                     |--S. baileyi (Prideaux & Wells 1998)LA02
                     |--S. brachyselenis (Prideaux & Wells 1997)LA02
                     |--S. browneorum (Merrilees 1968)LA02
                     |--S. cegsai (Pledge 1992)LA02
                     |--S. euryskaphus (Prideaux & Wells 1997)LA02
                     |--S. gilli (Merrilees 1965)LA02
                     |--S. maddocki (Wells & Murray 1979)LA02
                     |--S. mccoyi Turnbull, Lundelius & Tedford 1992LA02
                     |--S. newtonae Prideaux 2000LA02
                     |--S. oreas (De Vis 1895)LA02 [=Sthenurus (Simosthenurus) oreasF71]
                     |--S. orientalis (Tedford 1966)LA02 [=Sthenurus (Simosthenurus) orientalisF71]
                     |--S. pales (De Vis 1895)LA02 [=Sthenurus (Simosthenurus) palesF71]
                     `--S. tirarensis Prideaux 2004BA12

*Type species of generic name indicated


[BA12] Black, K. H., M. Archer, S. J. Hand & H. Godthelp. 2012. The rise of Australian marsupials: a synopsis of biostratigraphic, phylogenetic, palaeoecologic and palaeobiogeographic understanding. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 983–1078. Springer.

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[FS15] Faurby, S., & J.-C. Svenning. 2015. A species-level phylogeny of all extant and late Quaternary extinct mammals using a novel heuristic-hierarchical Bayesian approach. Molecular Phylogenetics and Evolution 84: 14–26.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[M62] Marcus, L. F. 1962. A new species of Sthenurus (Marsupialia, Macropodidae) from the Pleistocene of New South Wales. Records of the Australian Museum 25 (14): 299–304.

[PW10] Prideaux, G. J., & N. M. Warburton. 2010. An osteology-based appraisal of the phylogeny and evolution of kangaroos and wallabies (Macropodidae: Marsupialia). Zoological Journal of the Linnean Society 159 (4): 954–987.

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